Aplus transsylvanicus (Hoernes & Auinger, 1890)

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 35-36

Aplus transsylvanicus (Hoernes & Auinger, 1890)

Figs 15C , 18A–C

* Pollia multicostata Bell. var. transsylvanica —Hoernes & Auinger 1890: 236: pl. 28, figs 1–3.

Pollia multicostata Bellardi var. transsylvanica Hoernes et Auinger — Boettger 1906: 31.

Cantharus (Pollia) multicostatus var. transsylvanicus (Hoernes et Auinger, 1890) —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 173, pl. 43, figs 17a–b.

Cantharus (Pollia) multicostatus transsylvanicus (Hoernes & Auinger, 1890) — Atanacković 1985: 148, pl. 33, figs 11–12.

Aplus transsylvanicus (Hoernes et Auinger, 1890) — Kovács 2022: 90, figs 80–83.

Aplus transsylvanicus (Hoernes et Auinger, 1890) — Kovács & Vicián 2023: 251, figs 12F–G.

non Cantharus multicostatus transsylvanicus (Hoernes & Auinger, 1890) — Popa et al. 2014: 13, pl. 3, fig. 5 [= Polygona vindobonensis ( Csepreghy-Meznerics, 1956) ].

Type material. Lectotype designated herein: NHMW 1873 /0026/0019b, SL: 22.5 mm, MD: 11.5 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1890: pl. 28, fig 2), Figs 15C , 18A 1 –A 2 . Paralectotypes: NHMW 1873 /0026/0019a, SL: 24.3 mm , MD: 12.2 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1890: pl. 28, fig 1), 18B . NHMW 1873 /0026/0019b, SL: 19.4 mm , MD: 10.9 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1890: pl. 28, fig 3), Figs 18C 1 –C 2 .

Additional material. 2spec., NHMW1863 /0015/0437, Forchtenau( Austria) ; 46 spec., NHMW 1870 /0033/0087, Lăpugiu de Sus ( Romania) ; 1 spec., NHMW 1867 /0019/0085, CoŞteiu de Sus ( Austria) .

Revised description. Medium-sized, broad, ovate shell of seven teleoconch whorls; apical angle ~40°, increasing to ~60° on later spire whorls. Protoconch paucispiral of 1.5 convex whorls; diameter: 700 μm, height: 650 μm. Transition into teleoconch marked by onset of cancellate sculpture of prominent axial ribs, separated by interspaces of about equal width, overrun by three spiral cords, swollen over ribs. Later whorls with broad axial ribs, separated by narrower interspaces, overrun by prominent, convex spiral cords. Whorl profile of spire whorls weakly convex, periphery just below mid-whorl, suture deeply incised. Penultimate whorl evenly convex, sculpture increasing to eight spiral cords by bifurcation of adapical cord and intercalation of additional primary cords below mid-whorl. Last whorl attaining 65% of total height, regularly convex, moderately constricted at base, bearing 14–17 axial ribs, and about 16 raised, narrow spiral cords, interspaces with 1–2 weak secondary spiral threads, slightly scabrous due to raised growth lines; fasciole broad, indistinct with prominent, slightly scabrous spiral cords. Aperture wide, ovate. Columella broadly excavated, with six denticles decreasing in strength adapically. Lowermost two denticles most prominent, accentuating transition to siphonal canal. Columellar callus weakly thickened, sharply delimited from base, forming broad rim. Anal canal U-shaped, accentuated by small parietal denticle and more prominent anal denticle. Outer lip edge weakly crenulated, slightly thickened by weak terminal varix with about 11 prominent lirae from lip edge extending deep within aperture; lirae slightly thickened close to aperture. Siphonal canal moderately long, wide, only weakly deflected to the left, deeply notched.

Discussion. Aplus multicostata ( Bellardi, 1873) , from the Burdigalian or Langhian of the Colli Torinesi ( Italy), differs in its higher last whorl and the broader and more prominent axial ribs on the last whorl (see Bellardi 1873: pl. 12, fig. 15; Ferrero Mortara et al. 1981: pl. 4, fig. 10).

Paleoenvironment. Unknown, probably middle to outer neritic environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): Eisenstadt-Sopron Basin: Forchtenau ( Austria) (Hoernes & Auinger 1890); Pannonian Basin: Letkés ( Hungary) ( Kovács & Vicián 2023); Southern Pannonian Basin: Hrvaćani ( Bosnia and Herzegovina) ( Atanacković 1985); Făget Basin: CoŞteiu de Sus, Lăpugiu de Sus ( Romania) ( Kovács 2022).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 15. Protoconchs of Paratethyan Pisaniidae, Prodotiidae and Tudiclidae. A. Aplus moravicus (Hoernes & Auinger, 1890), NHMW 1862/0029/0034, lectotype, Rudice u Blanska (Czech Republic). B. Aplus subpusillus (Hoernes & Auinger, 1890), NHMW 1866/0001/1066, paralectotype, Forchtenau (Austria). C. Aplus transsylvanicus (Hoernes & Auinger, 1890), NHMW 1873/0026/0019b, lectotype. D. Aplus zebus (De Gregorio, 1885a), NHMW 1855/0045/0439, Gainfarn (Austria). E. Aplus anatolicus nov. sp. RGM 784 005, paratype, Seyithasan (Turkey). F. Hilda transsylvanica (Hoernes & Auinger, 1884), NHMW 1876/0011/0023a, lectotype, Lăpugiu de Sus (Romania).G. Janiopsis vindobonensis nov. sp., NHMW 1850/0009/0002, holotype, Baden (Austria). H. Monostiolum hungaricum (Csepreghy-Meznerics, 1954), NHMW 1868/0001/0094, Steinebrunn (Austria). I. Dianthiphos klaudiae (Kovács & Vicián, 2023), NHMW1846/0037/0144a, Steinebrunn (Austria). J. Euthria obelixi nov. sp., NHMW 1865/0015/0060, holotype, Lysice (Czech Republic). K. Tethyspollia mariae (Hoernes & Auinger, 1890), NHMW 1868/0001/0441, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 18. Aplus transsylvanicus (Hoernes & Auinger, 1890). A1–A2. NHMW 1873/0026/0019b, lectotype, Lăpugiu de Sus (Romania). B. NHMW 1873/0026/0019a, paralectotype, Lăpugiu de Sus (Romania). C1–C2. NHMW 1873/0026/0019b, paralectotype, Lăpugiu de Sus (Romania).