Aplus ranellaeformis (Hoernes & Auinger, 1890)

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 33-34

Aplus ranellaeformis (Hoernes & Auinger, 1890)

Figs 16A–B

* Pollia ranellaeformis nov. form.—Hoernes & Auinger 1890: 238, pl. 28, figs 10a–b.

Pollia ranelliformis [sic] Hö. Au.— Boettger 1902: 34.

Janiopsis ranelliformis [sic] (Hö. Au.)— Boettger 1906: 32.

Janiopsis labrosa var. orientalis n. var. —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 175, pl. 44, figs 3–4.

Janiopsis labrosa ( Bellardi et Michelotti, 1840) — Kovács 2022: 91, figs 86–89 [non Janiopsis labrosa ( Bellardi & Michelotti, 1840) ].

Type material. Lectotype (designated herein): NHMW 1867 /0019/0102, SL: 14.7 mm, MD: 8.1 mm, CoŞteiu de Sus ( Romania), illustrated in Hoernes & Auinger (1890: pl. 28, fig. 10), Figs 16A 1 –A 2 . Paralectotype: NHMW 1867 /0019/0104, SL: 16.3 mm , MD: 9.2 mm, CoŞteiu de Sus ( Romania), Figs 16B 1 –B 2 .

Revised description. Small, moderately broad shell. Protoconch and early teleoconch whorls unknown. Teleoconch whorls with shallow, slightly convex subsutural ramp, weakly convex periphery, angled just above incised suture.Sculpture of broad axial ribs, separated by interspaces of about equal width, overrun by three prominent spiral cords on first teleoconch whorl, abapical cord most prominent, with single slightly weaker secondary cord intercalated in interspaces; cords swollen over ribs; nine spiral cords on penultimate whorl. Last whorl attaining ~78% of total height, weakly shouldered, bearing ten axial ribs and strong terminal varix; base moderately constricted bearing prominent spirals of alternating strength; fasciole broad, indistinct with three raised primary spiral cords separated by weaker secondaries. Aperture moderately wide, ovate. Columella strongly excavated in upper half with three blunt denticles on abapical half; lowermost delimiting siphonal canal. Columellar callus forming broad rim, moderately delimited from base. Anal canal U-shaped, moderately incised, accentuated by weak parietal denticle and prominent anal denticle. Outer lip thick with about six prominent, elongated denticles (or short lirae). Siphonal canal moderately short, wide, slightly deflected to the left, shallowly notched.

Paratethyan synonyms. Janiopsis labrosa orientalis Kojumdgieva in Kojumdgieva & Strachimirov, 1960, from the Badenian of Bulgaria, is most probably a subjective junior synonym of Aplus ranellaeformis . Unfortunately, it was not possible to trace the types of the Bulgarian species.

Discussion. Aplus nilus De Gregorio, 1884 , from the Pliocene of Italy, has a comparable outline but differs in its less prominent axial ribs, broader primary spiral cords and weaker terminal varix (see Brunetti & Della Bella 2014: figs 2B–G, 3A–D). Aplus dispar ( Millet, 1865) , from the Tortonian of the Loire Basin ( France), develops a similar sculpture and has angled whorls but has much narrower axial ribs and a weaker terminal varix (see Landau et al. 2019: pl. 43). ‘ Janiopsis ’ labrosa ( Bellardi & Michelotti, 1840) , from the Early Miocene of the Colli Torinesi ( Italy), does not have the strong parietal and labial teeth flanking the anal canal seen in Aplus ranellaeformis (Hoernes & Auinger, 1890) .

Paleoenvironment. Unknown. Distribution in Central Paratethys. Badenian (Middle Miocene): Făget Basin: CoŞteiu de Sus, Lăpugiu de

Sus ( Romania) ( Boettger 1906; Kovács 2022; hoc opus); Dacian Basin: Opanec, Staropatica, Târnene ( Bulgaria)

(Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 16. Aplus ranellaeformis (Hoernes & Auinger, 1890). A1–A2. NHMW 1867/0019/0102, lectotype, CoŞteiu de Sus (Romania). B1–B2. NHMW 1867/0019/0104, paralectotype, CoŞteiu de Sus (Romania). C. Aplus volhynicus (Friedberg, 1912), lectotype, Nr. 92a, State Natural History Museum, National Academy of Sciences of Ukraine, Żabiak (Ukraine).