Phasia obesa ( Fabricius, 1798 )

Sun, Xuekui & Marshall, Stephen A., 2003, Systematics of Phasia Latreille (Diptera: Tachinidae), Zootaxa 276 (1), pp. 1-320 : 92-97

3.4.13 Phasia obesa ( Fabricius, 1798)

(Figures I­43, II­11.3)

Thereva obesa Fabricius 1798: 561 .

Musca nebulosa Panzer 1798: 20 .

Thereva cinerea Fabricius, 1805: 221 .

Thereva muscaria Fallén, 1851: 232 .

Phasia atropurpurea Meigen, 1821: 193 .

Phasia violacea Meigen, 1824: 193 .

Phasia hamata Meigen, 1824: 193 .

Phasia albipennis Meigen, 1824: 194 .

Phasia nubeculosa Meigen, 1824: 195 .

Phasia umbripennis Meigen, 1824: 195 .

Phasia nervosa Meigen, 1824: 196 .

Hyalomyia nebulosa Robineau­Desvoidy, 1830: 300 .

Hyalomyia fuscipennis Macquart, 1835: 203 .

Phasia speciosa Curtis, 1838: 697 .

Phasia (Alophora) aenea von Roser, 1840: 57 .

Phasia umbrata Zetterstedt, 1844: 1249 .

Phasia grisea Zetterstedt, 1844: 1250 .

Phasia flavipennis Zetterstedt, 1844: 1250 .

Hyalomyia fuligipennis Rondani, 1861: 215 .

Hyalomyia murina Rondani, 1861: 216 .

Hyalomyia areolaris Rondani, 1861: 216 .

Hyalomyia purpurea Robineau­Desvoidy, 1863: 244 .

Hyalomyia coerulescens Robineau­Desvoidy, 1863: 245 .

Hyalomyia nitida Robineau­Desvoidy, 1863: 250 .

Hyalomyia fuscana Robineau­Desvoidy, 1863: 254 .

Hyalomyia atra Robineau­Desvoidy, 1863: 254 .

Hyalomyia tomentosa Rondani, 1868: 603 .

Hyalomyia umbripennis Girschner, 1886: 5 (as Hyalomyia obesa var.).

Hyalomyia nebulosa Girschner, 1886: 65 (as Hyalomyia obesa var.).

Hyalomyia fascipennis Girschner, 1886: 66 (as Hyalomyia obesa var.).

Hyalomyia latipennis Girschner, 1886: 68 (as Hyalomyia obesa var.)

Alophora (Hyalomyia) obesa: Girschner, 1886: 1 ­ Girschner, 1887: 28 ­ Bezzi et Stein 1907: 580 ­ Lundbeck 1927: 95 ­ Wainwright 1928: 244 ­ Emden 1954: 27 ­ Rohdendorf 1947: 86 ­ Day 1948: 41.

Allophora obesa: Baer 1921: 128 .

Alophora obesa: Stein 1924: 259 ­ Rohdendorf 1933: 712 ­ Belanovskij 1951: 140.

Alophorella obesa: Townsend 1938: 36 .

Allophora (Allophorella) obesa: Dupuis 1949: 505 .

Allophorella obesa: Dupuis 1963: 105 (host, catalog).

Alophora (Alophorella) obesa: Draber­Mońko 1965: 159 .

Phasia (Phasia) obesa: Herting 1984: 169 (catalog) ­ Herting and Dely­Draskovits 1993: 410 (catalog).

DESCRIPTION

Body length: 6­8 mm.

MALE: HEAD. Head spherical or nearly so. Eyes separated by a distance wider than ocellus but narrower than ocellar triangle. Fronto­orbital plate black, yellow or grey pruinose, 4­5 rows of hairs laterally. Frontal vitta black. Frontal vitta at base of antennae 0.8 times as wide as fronto­orbital plate anteriorly. Ocellar setae present but fine; outer vertical setae present; inner vertical setae absent. Face black with grey pruinosity. Parafacial black with grey pruinosity, bare, 1.1 times as wide as first flagellomere. Lower margin of face projecting, visible in profile, brown to black with grey pruinosity. Vibrissa well differentiated; intervibrissal distance 1.1­1.2 times distance between vibrissa and eye on same side; facial ridge with bristles on lower 1/4. Gena brown to black with grey pruinosity; hairs white; height 0.12 times eye height. Lunule black and shining; sublunular bulla indistinct. Antenna pedicel yellow or brown, first flagellomere black, 1.2 times as long as pedicel; arista thickened on basal 0.4. Length of oral opening 3.1 times its width. Occiput flattened, white pruinose; hairs white. Palpus yellow, or brown.

THORAX. Mesoscutum strongly pruinose; 4 black longitudinal vittae, with fine black hairs. 0+1 acrostichal seta; 0+1 dorsocentral seta; 1­2 postpronotal setae; presutural supraalar seta present, strong; 1 postsutural intra­alar seta; 2 notopleural setae; 1 supra­alar seta; 2 postalar setae. Pleuron thinly grey pruinose; hairs black and brown; and without scalelike setae. Anepimeral setae medium size, black. 1 katepisternal seta. 4­9 meral setae. Scutellum black with grey pruinosity, with two pairs of marginal setae; apical setae present and distinct; discal setae randomly arranged, not in rows. Subscutellum not very prominent. WING. Lower calypter hyaline white. Wing base without scale­like setae. Tegula black. Basicosta brown to black. Wing hyaline or pictured, narrow or broad; petiole of apical cell 0.20­0.24 times as long as preceding section of R 4+5; M meeting R 4+5 almost at right angle. Halter yellow, or brown. LEGS. Fore femur swollen black; ventrally and laterally with whitish yellow hairs, dorsally with black hairs. Fore tibia black, without bristles, apically with 1 pv and 1 d. Fore tarus black. Fore claws brown; 1.1­1.3 times as long as fifth tarsomere. Pulvilli brown. Mid femur black, with whitish yellow hairs, or hairs black. Mid tibia brown to black, with 0­1 ad, apically with 1 ad, 1 a, 1 av, 1 pv, 0­1 p, 1 v and 1 d. Hind femur brown to black; whitish yellow hairs, or hairs black; scale­like setae absent. Hind tibia brown to black, 6 ad and 3­4 pd.

ABDOMEN. Abdominal tergites black; longitudinal vitta absent or present; dorsum of tergites purple shiny, or from thinly to strongly grey pruinose. Relative length of abdominal tergites I+II:III:IV:V = 0.4:1:1:0.7; and relative width I+II:III:IV:V = 1.5:1:1:1.2. Hair spots indistinct. TERMINALIA. Syncercus with a semicircular notch posteriorly; Surstylus slender, longer than cercus. Ejaculatory apodeme small, knob­like. Epiphallus well developed. Pregonite long, pointed, longer than postgonite, haired dorsally. Postgonite pointed. Phallus and phallapodeme longer than hypandrium. Phallus densely haired dorsally. Distiphallus not well sclerotized.

FEMALE. Wing hyaline. Eyes separated by a distance as wide as single ocellus. Anterior spine­like setae of hind tibia present. Sternite VII (sheath) longer than sternite VI; tapered, narrow apically; not bent, apex straight; sternite VII with linear wrinkles ventrally. Ovipositor bent upward.

TYPE MATERIAL

Thereva obesa Fabricius , 1798.0 Holotype ♂, Italy (Copenhagen or lost, Townsend 1938: 36, not examined).

Hyalomyia obesa umbripennis Girschner, 1886 . Syntypes, 2♂, "Ex: Girschner Collection" [no data] (BMNH, examined).

Hyalomyia obesa nebulosa Girschner, 1886 . Syntypes, 1♂, "Ex: Girschner Collection" [no data] (BMNH, examined).

Hyalomyia obesa fascipennis Girschner, 1886 . Syntypes, 2♂, "Ex: Girschner Collection" [no data] (BMNH, examined).

Hyalomyia obesa latipennis Girschner, 1886 . Syntypes, 2♂, [ SWITZERLAND], Meirngeu?, ex: Girschner Collection, [no data] (BMNH, examined).

OTHER MATERIAL EXAMINED

AUSTRIA. Matrei in Osttirol, 14.viii.1967 (1♂, BMNH); Raidina and Unterfravenhaid, vii­viii.1980, ex: Lygus sp. , H. Hoyer (1♂ 1♀, USNM); Wien, viii.1940, H. Loew (1♂, ZMHU) Feldkirch, Vorarlberg, Moosbrugger (1♀, DEIC); Zillertal, umg. Mayrhofen, 4.vii.1914, H. Wagner (1♂ 4♀, DEIC). CHINA. Inner Mongolia, Ulan Qi, 4.viii.1971 (1♂ 1♀, IZAS); Sichuan [Szechuen], Yellow Dragon Gorge, near Sougpan 1924, 12,000 ­ 14,000 ft, D.C. Graham Coll.(1♂, USNM); Sichuan, Yanyuan, 2,200m, 18.ix.1986, Y. Wang (1♂, IZAS); Tibet, 10,000ft, Yatung, viii.1927, Bailey (1♀, BMNH); Tibet, near Tang­Gu, 14,000ft, 3­6.viii.1930, D.C. Graham (1♂, USNM); Penniu, 28.ix.1925, C.Y. Wong (1♀, USNM). CROATIA. Elviacliu (?), 19.viii. 1933, N. Baranov Coll.1960 (1♂, USNM); Pcetenica, 19.vii.1016 (1♂ 1♀, USNM); Twcopalze (?), 25.vi.1916, N. Baranov Coll.1960, (1♂, USNM); Zagreb, 25.vii. 1908, 20.vii.1929 and 18.ix. 1929, N. Baranov Coll. 1960 (3♂ 1♀, USNM). CYPRUS. Amathus, 14.ii.1936, G.A. Mavromoustakis (1♂, BMNH); C. Glaszner (2♂, BMNH); 12.v.1928, H.M. Morris (1♂, BMNH); Dnavlos, 10.iv.1971, K. Guichard (1♂, BMNH); Kantara, 18.v.1948, G.A. Mavromoustakis (7♂, BMNH); Mt. Troodos, 1.vii.1937, G.A. Mavromoustakis (2♂, BMNH). CZECHOSLOVAKIA. Carlsbad, 8­22.viii.1915, coll. Oldenberg (3♂ 4♀, DEIC). Bohemia, mer. Horusicky, Rybnik, 4.9.1967, M. Chvala (1♀, BMNH). DENMARK. Nielsen, W.R. Walton Collection (1♂, USNM). ENGLAND. Jersey, St. Catharines, 6.8.46, (1♂, USNM). FRANCE. Ardennes, Vendresse, 17.vii.1912 (1♂, USNM); Argeles­Gazost, South France, 10.viii.1911, C.J. Wainwright (1♀, BMNH); Colmar, H.R., ex: Hypera postica , 24.vi.1964, J. Balakine (1♀ with pupa, USNM); Fontainbleau (21km N), ex: Lygus sp. 29.viii.1962, R.I. Sailer (1♂ with pupa, USNM); Hautes Alpes, Brunissard, 5.vii.1992, Tschorsnig (1♂, DEBU); La Gueue les Yuelines, 15.viii.1936, (1♀, CNCI); Marans (M.­et­ L.), 10­7­21, (1♀, BMNH); Marans (M.et L.), 4.vii. 1921 (3♂, BMNH); Richelieu, 27.viii and 13.ix.1965, C. Dupuis (3♂, USNM); Sacias, 10.viii.1927 (1♀, BMNH); Vernet les Bains, Pyrénées­Or., 15.vi.1920, C.J. Wainwright (1♀, BMNH); Void, M.and M. ex: Lygus rugulipennis , 5.v.1963 (adults) and 20.v.1963, R.I. Sailer (1♂ 1♀, with pupae, USNM). GERMANY. Berlin, 1894­1912, (31♂ 19♀, DEIC); Berlin, 4.viii.1926, G. Enderlein (1♂, ZMHU); Dresden, 21.viii.1903, coll. Liehtwardt (11♂ 5♀, DEIC); Fùrth, I.B. Dr. Trautmann, 1918, David, G. Hall Coll. (2♂ 1♀, USNM); Frankfurt Oder, M.P. Riedel, Reinhard Collection (6♂ 4♀, CNCI); Potsdam, 6.vi.1916, 27.vi.1909 (5♂, DEIC); Rangsdorf, 10.8.1918, coll. Oldenberg (26♂ 2♀, DEIC); Sachsen, Genthin, 20.vii.1895, P. Stein (1♂, ZMHU); Riedel Pössneck, coll. Oldenberg (3♂ 3♀, DEIC); Thal, i. Th, viii.1894, coll. Oldenberg (5♂ 2♀, DEIC). GREECE. Creta, v.0. coll. Liehtwardt (6♂, DEIC). HUNGARY. Budapest, 21.vi.1901, coll. Liehtwardt (1♀, DEIC); Szár, 15.vi.1902, coll. Liehtwardt (4♂ 1♀, DEIC). INDIA. Tribeni, 6.vi.1943, H. Loew (1♀, ZMHU); Pcuni, Grunei, L. Loew (1♂, ZMHU). ISRAEL. Latroun, 22.xi.1968, S. Bleszynski (1♂, CNCI). ITALY. [?], (1♀, DEIC); Bozen, vi.1996, coll. Oldenberg (1♀, DEIC); Eyrs, 5.8.09, coll. Oldenberg (1♂, DEIC); Macugnaga, 21.vii.1900, Oldenberg (2♂ 1♀, DEIC); Lia Centrale, 28.x.1918 (1♂, BMNH); Umbria Prov., Trevl., 13.viii.1944, G.E. Shewell (1♀, CNCI); Villalatina (Prov. Caserta), 1,500ft, O. Querci (1♂, BMNH); Marche, Monti Sibillini, Bolognola, 1­20.vii.1935, L. Ceresa (1♂, AMNH). JAPAN. Kyushu, Mt., Tachibana, Fukuoka City, 8.v.1980, H. Shima (1♀, BLKU); Obihiro, S. Takano (1♂, CNCI). LATVIA. Curland, C. Siebert (8♂ 12♀, DEIC). LEBANON. Beirut, Amchit, 35 Km N. 23­30.ix.1971, A.K. Walker (8♂ 10♀, CNCI). MOROCCO. Asni, viii.1930, A. Mackie (1♀, BMNH). NORWAY. Opland, Breivegen Bru., 4­18.viii.1979, S. Andersen (1♂, CNCI); Eidskog, ix.1900, N. Strand (5♂ 3♀, DEIC), POLAND. Bogucice, 21.vii.1963, A. Mońko (1♀, USNM); Bieszczady, 23.ix.1968, S. Bal (1♂, USNM); Schlesien, coll. Letzner (9♂ 4♀, DEIC); Urocz, Sokotöh D., 7.vii.1961, A. Mońko (1♂, USNM). RUSSIA [USSR]. Siberia, Novosibirsk Region, Chany Lake Biological Station, near Zdvensk, on flowers, 1­5.vii.1991, S.A. Marshall (8♂ 7♀, DEBU, LACM); Tolmatschevo, vic. Luga, 17.vii.1937, and 30.vii.1936, Stackelberg (2♀, USNM). SPAIN. Lerida, Val d'Aran, Valencia de Aneu, 18.ix.1956 (1♀, BMNH); Merrill, H. Prime 6.viii.1957 (1♂, CNCI); Prov. Salamanca, Villar de Ciervo, 20.v.1988 Tschorsnig (1♀, DEBU); Prov. Lérida, Vallar de Aran, Arties, 29.vi.1992, Tschorsnig (1♂, DEBU); Prov. Lérida, 6 km NW Llavorsi, 1.vii.1992, Tschorsnig (1♂, DEBU); Prov. Huesca, Linás de Broto, 26.vi.1992, Tschorsnig (1♂, DEBU). SWEDEN. Bolka, SE of Oregrund, 14.vii.1980, A.C. Pont (1♂,BMNH). SWITZERLAND. Delémont, 900m, 30­31.vii.1958, H. Pschorn (1♂ 1♀, CNCI); Delémont, 14­24.viii.1964, C.W. Sabrosky (1♂, USNM); Delémont, Domont, 26.viii.1964,(1♂, DEBU); Delémont, Domont, 13­23.ix.1966 (6♂, BMNH); Delémont, 8­23.viii.1971 ­74, 28.viii.1964 (6♂ 2♀, CNCI); Eclepens, 10.viii.1914. C.J. Wainwright Collection, B.M.1948­488 (1♀, BMNH); Meirngeu[?], ex: Girschner Collection, C.J. Wainwright Collection, B.M. 1948­488 (93♂, BMNH). YOGOSLAVIA. Kcanjska Gola, 30.v.81, P.M. Ackland (1♂, BMNH). DOUBTFUL LOCALITIES. "Beths, sunninghill, 20.vii.1948 (1♀, USNM)"; "Achau, David G. Hall Coll. (1♀, USNM)"; "Muauof(?), 29.vi.1916 (1♂, USNM)"; "Kis­Pöse, Méhely (1♂, USNM)"; "A.L. Melander Collection 1961, with det label by L. Strobl (2♂, USNM)"; "Europe: (no detailed location data or unreadable) (51♂ 21♀, DEIC)"; "Tullu­A, ex: Lygus rugulipennis , 15.viii.1977, L.P. Mesnil (1♂ 1♀, CNCI)"; "Dania, B. VB92, Slotslyngen, 30.vi. 10.vii.1981, S. Andersen (1♀, CNCI)"; "Ingeiheimsche Au. b. Geisenheim, 19.v.1923 (1♀, CNCI)"; "Geisenheim (Rheingau), 23.ix.1922 (1♂, CNCI)"; "Riedel Possneck, Reinhard Collection (1♂ 1♀, CNCI)"; "ex: Girschner Collection, C.J. Wainwright Collection, B.M. 1948­488,(no other readable data) (7♂ 1♀, BMNH)"; " ex: Girschner Collection (45♂ 45♀, BMNH)".

HOSTS

Zicrona caerulea (L.) ( Hemiptera , Pentatomidae ): ­ (Dupuis 1949: 505, 1963: 105 ­ Draber­Mońko 1965: 78).

Leptopterna dolabrata (L.) ( Hemiptera , Miridae ): ­ (Southwood and Leston 1959: 313 ­ Dupuis 1963: 105 ­ Draber­Mońko 1965: 78).

Lygus pratensis (L.) ( Hemiptera , Miridae ): ­ ( Dupuis 1963: 105 ­ Draber­Mońko 1965: 78).

Myrmus miriformis (Fallen) ( Hemiptera , Rhopalidae ): ­ ( Dupuis 1963: 105 ­ Draber­ Mońko 1965: 78).

Beosus maritimus (Scopoli) ( Hemiptera , Lygaeidae ): ­ ( Dupuis 1963: 105 ­ Draber­ Mońko 1965: 78).

Lygus rugulipennis Poppins ( Hemiptera , Miridae ) ­ France.

Lygus sp. , ( Hemiptera , Miridae ): ­ France (Fontainbleau), Austria (Raidina and Underfravenhaid).

Doubtful Host. Hypera postica (Gyllenhal) ( Coleoptera , Curculionidae ) ­ France (Colmar).

NOTES

Phasia obesa (Fabricius) is highly variable, especially for wing patterns, abdominal pruinosity and the shape of sheath (see Draber­Mońko 1965: 159­167 [figs: 322­357]).

This species is similar to the Nearctic species Phasia aeneoventris (Williston) and Phasia robertsonii (Townsend) , and the Neotropical species Phasia moerens (Wulp) . Males of Phasia obesa can be separated from these species on the basis of the distiphallus and pregonite, but no diagnostic character has been found for females.