Smicromyrmini Bischoff, 1920

Lelej, Arkady S. & Williams, Kevin A., 2023, Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae), Zootaxa 5231 (2), pp. 101-140 : 103-104

Tribe Smicromyrmini Bischoff, 1920

Smicromyrmini Bischoff, 1920: 23 ; Invrea 1964: 151 (Smicromyrminae); Brothers 1975: 592 (Smicromyrmina); Lelej 1985: 180; Lelej 1995b: 2; Lelej & Nemkov 1997: 4; Brothers 1999: 245 (Smicromyrmina); Brothers & Lelej 2017: 47; Pagliano et al. 2020: 169.

Diagnosis. MALE. Eye without setae; inner margin deeply notched. F1 shorter than F2. Scutellum simple, without median carina or postero-lateral denticle. Wing venations not reaching outer wing margin; distance between origin of RS on vein Sc and base of stigmatic cell nearly 2 × stigmatic cell length; hind wing without anal lobe. Tarsal claws without inner denticles. Metasomal segment 1 narrow; segment 2 with lateral felt line on T2 and sometimes on S2; S8 not reduced, plate-like; genitalia with symmetrical penis valves. FEMALE. Head not enlarged; eye oval, slightly protruding from head capsule, without setae; maxillary palpus six segmented; labial palpus four segmented. Mesosoma widest at pronotum or propodeum; mesopleuron concave; meso-metapleural suture ending at pronotal spiracle; dorsum of propodeum without row of denticles. Claws without inner denticles. T1 much narrower than T2, not petiolate; T2 with one or three, (two lateral transverse in some Mickelomyrme , subbasal band in some Dentilla , discal spots entirely obliterated in some species of many genera) subbasal spots of pale setae or without any spots.

Genera included. 26 genera and 5 subgenera ( Brothers & Lelej 2017, Pagliano et al. 2020).

Diversity and distribution. Widely distributed in the Old World and numbers 26 genera, 528 species:Palaearctic (13/213), Afrotropical (15/126), and Oriental (14/204) Regions (Lelej 2007, Pagliano et al. 2020, Thaochan et al. 2022). The endemic genera are: Nuristanilla Lelej in Lelej & Kabakov, 1980, ♀ and Skorikovia Ovtchinnikov, 2002 , ♁, ♀ in the Palaearctic Region; Indratilla Lelej, 1993 , ♁, ♀; Karunaratnea Lelej, 2005 , ♁, ♀; Nordeniella Lelej, 2005 , ♁, ♀, and Tsunekimyrme Lelej, 1995b , ♁ in the Oriental Region; and Antennotilla Bischoff, 1920 , ♁; Corytilla Arnold, 1956 , ♁, ♀; Ctenoceraea Nonveiller, 1993 , ♁; Guineomutilla Suárez, 1977 , ♀; Gynandrotilla Arnold, 1946 , ♁; Paglianotilla Lelej in Lelej & van Harten, 2006, ♁; Psammotherma Latreille, 1825 , ♁; Pseudocephalotilla Bischoff, 1920 , ♁, ♀; and Sulcotilla Bischoff, 1920 , ♁, ♀ in the Afrotropical Region.

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