Quadraceps separatus ( Kellogg & Kuwana, 1902 )

Palma, Ricardo L. & Peck, Stewart B., 2013, <strong> An annotated checklist of parasitic lice (Insecta: Phthiraptera) from the Galápagos Islands </ strong>, Zootaxa 3627 (1), pp. 1-87 : 55-56

Quadraceps separatus ( Kellogg & Kuwana, 1902)

Nirmus separatus Kellogg & Kuwana, 1902: 472 , pl. 29, fig. 6.

Nirmus gloriosus emarginatus Kellogg & Chapman, 1902: 159 .

Nirmus separatus Kellogg & Kuwana, 1902 : Kellogg 1906: 317. In part Nirmus separatus ; in part Philoceanus becki Kellogg, 1903 .

“ Lipeurus helleri ” Kellogg, 1906: 319 (not Lipeurus helleri Kellogg & Kuwana, 1902 ). In part Quadraceps separatus ; in part Pectinopygus minor ( Ewing, 1924) ; in part Quadraceps birostris ( Giebel, 1874) .

Degeeriella emarginata ( Kellogg & Chapman, 1902) ; Harrison 1916: 112.

Degeeriella separata ( Kellogg & Kuwana, 1902) ; Harrison 1916: 123.

Quadraceps separatus ( Kellogg & Kuwana, 1902) ; Hopkins & Clay 1952: 316

Quadraceps separatus ( Kellogg & Kuwana, 1902) ; Clay in Linsley & Usinger 1966: 131.

Anousceps separatus ( Kellogg & Kuwana, 1902) ; Złotorzycka 1967: 731.

Quadraceps separatus ( Kellogg & Kuwana, 1902) ; Palma 1995: 221.

Lectotype ♂ in EMEC ( Palma 1996: 215).

Type host: Geospiza conirostris Ridgway, 1890 , in error (see Palma 1994b: 270).

Galápagos host: Anous stolidus galapagensis Sharpe, 1879 .

Galápagos localities: Isla Wolf; Isla Marchena; Isla Seymour; Isla Rábida.

Galápagos references: Kellogg & Kuwana (1902); Kellogg (1906); Kellogg (1908: 29); Thompson (1939: 121); Linsley & Usinger (1966); Palma (1994b: 268; 1995).

Other significant references: Ferris (1932: 69, figs 19a–e); Timmermann (1952b: 73, figs 1–2; 1957: pl. 9, figs c–d); Amerson & Emerson (1971: 18, 25); Palma (1996: 215); Price et al. (2003: 227).

Material examined: 90 males, 89 females and 10 nymphs (6 samples, EMEC, MONZ) .

Remarks: From the Kellogg Collection, we examined the lectotype of Nirmus separatus from Geospiza conirostris , and seven paralectotypes all associated with incorrect hosts (see Palma 1994b: 270). Also, we examined five specimens labelled and reported by Kellogg (1906: 319) as “ Lipeurus helleri ” and three specimens labelled and reported by Kellogg (1906: 317) as “ Nirmus separatus ”, all from Anous stolidus galapagensis , and have identified them as Quadraceps separatus . At present, Anous stolidus is the only host of this louse species ( Price et al. 2003: 227).