Myosoma lagopus (KRIECHBAUMER), 2012

Papp, J., 2012, A Taxonomic Study Of The Myosoma Genus-Group With Description Of Amyosoma Cavei Sp. N. From Honduras (Hymenoptera: Braconidae: Braconinae: Braconini), Acta Zoologica Academiae Scientiarum Hungaricae 58 (1), pp. 1-29 : 20-22

Myosoma lagopus (KRIECHBAUMER) sp. rev. et comb. n.

( Figs 74–83 )

Acanthobracon lagopus KRIECHBAUMER, 1900: 103 ♀ (1 ♀), type locality: “Zwischen Bodega central und Honda am Rio Magdalena (Columbien)”, female holotype (designated by “Zoologische Staatssammlung München”) in Museum München; examined. – SCHULZ 1903: 253 (synonymized with M. hirtipes BRULLÉ, 1846 ). SHENEFELT 1978: 1708 (as synonym of M. hirtipes after SCHULZ l.c., literature up to 1903).

Bracon errotus SZÉPLIGETI, 1902: 43 ♀ (1 ♀), type locality: “ Venezuela: Merida ”, female holotype (designated by J. PAPP 1969 in QUICKE 1991: 172) in Magyar Természettudományi Múzeum, Budapest; examined, syn. n. – SZÉPLIGETI 1904: 187 (in key).

Myosoma errotus (SZÉPLIGETI) : SZÉPLIGETI 1906: 588 (comb. n.). SHENEFELT 1978: 1708 (as valid species, literature up to 1906).

Designation of the female holotype of Acanthobracon lagopus – (First label, handwriting) “Bodega central de Honda”; (second label, handwriting) “Bodega central / und Honda / am Rio Magda- / lena, Columbien / Therese v. Bayern”; (third label, reverse second label) “Bisher war nur des ♂ / n. als Fundort / «Amer. mer.» bekannt”; fourth label is the holotype card, fifth and sixth labels are with the valid (“recte”) name Myosoma hirtipes Brullé ♀ (the two labels are with two different handwriting). – Holotype is in fairly good condition: (1) pinned by mesosoma; (2) head glued to mesosoma; (3) metasoma, right hind coxa and right hind leg glued separately on hind part of first label; (4) right antenna missing; (5) right fore tibia less visible (glued to head).

Designation of the female holotype of Bracon errotus – (First label, my handwriting) “ Venezuela / Merida ”; (second quadratic label, reverse the first label) “539/128”; (third label) “ Br. errotus ” (Szépligeti’s handwriting) / “det. Szépligeti” (printed); (fourth label) “ Myosoma errotus (Szépl.) ♀ (my handwriting) “det. Szépligeti” (printed) “1906” (my handwriting); fifth label is the holotype card, sixth label is with the inventory number “1248”; seventh label is with the actual name Myosoma lagopus KRIECHBAUMER (det. J. Papp 2009), specimen compared with the female holotype of M. lagopus . Labels 1 and 4 to 7 attached by me. – Holotype is in good condition: (1) pinned by mesosoma (before prescutellar furrow); (2) left flagellum damaged; (3) right fore wing damaged proximally from pterostigma.

Redescription of the female holotype of Acanthobracon lagopus – Body 8.5 mm long. Left antenna about as long as body and with 49 antennomeres (right antenna missing). Scape in outer-lateral view 1.5 times as long dorsally as broad apically, dorsally longer than ventrally ( Fig. 74 ). Flagellomeres 1–3 1.25 times, 1.08 times 1 times as long as broad, respectively, further flagellomeres transverse (i.e. slightly broader than long), penultimate three flagellomeres cubic. Flagellum distally indistinctly attenuating, hairy. – Head in dorsal view less transverse ( Fig. 75 ), 1.6 times as broad as long, eye 1.6 times longer than temple, tempel receded. Ocelli middle-sized, elliptic, OOL slightly more than three times longer than POL ( Fig. 75 ). Eye in lateral view 1.5 times as high as wide, temple beyond eye 0.7 times as wide as eye and narowing ventrally ( Fig. 76 , see arrows). Horizontal diameter of oral opening 2.6 times longer than shortest distance between opening and eye. Head polished, face weakly uneven.

Mesosoma in lateral view 1.5 times as long as high, polished. Notaulix distinct, smooth. – Hind femur 3.3 times as long as broad medially and with fairly long hairs ( Fig. 77 ). Hind basitarsus as long as tarsomeres 2–4 combined. Claw downcurved and with wide and somewhat pointed basal lobe ( Fig. 78 ).

Fore wing as long as body. Pterostigma ( Fig. 79 ) 3.3 times as long as wide and issuing r from its middle, r as long as width of pterostigma. Second submarginal cell short, 3–SR 1.6 times length of 2–SR; SR1 almost straight, reaching tip of wing and 1.7 times longer than 3–SR. First discal cell: 1–SR–M weakly bent and almost twice longer than 1–M, 1–M and m–cu not parallel ( Fig. 80 ). – Hind wing: 1r–m curved like that of M. hirtipes (cf. Fig. 73 ).

First tergite ( Fig. 81 ) five times as long as broad, scutum moderately broadening posteriorly, together with further tergites polished. Second tergite transverse and laterally narrowing, four times broader behind than long medially; third tergite as long as second tergite ( Fig. 81 ). Ovipositor sheath long, as long as hind tibia + half basitarsus combined.

Antenna, had, mesosoma, legs and ovipositor sheath black, metasoma reddish. Mandible dark rusty. Upper margin of pronotum rusty. Hairs of legs black. Wings brown fumous, pterostigma and veins brown.

Deviating features of one female (holotype of M. errotus ).–Body 8 mm long. Head in dorsal view 1.5 times as broad as long ( Fig. 82 ). Hind femur four times as long as broad medially ( Fig. 83 ). Fore wing: 3–SR 1.5 times as long as 2–SR, SR1 1.6 times longer than 3–SR. Outer orbit (of eye) rusty, pronotum and lateral lobe of mesoscutum with faint rusty suffusion.

Male and host unknown.

Distribution – Colombia, Venezuela.

Taxonomic position – Since SCHULZ’ s synonymization (l.c.) the name A. lagopus was suppressed as conspecific with M. hirtipes ( SHENEFELT 1978) . The examination of the holotypes of the two taxa, M. hirtipes and M. lagopus albeit very near to each other, shows two valid species. Their distinction is presented as follows:

1 (2) Second submarginal cell long, 3–SR twice as long as 2–SR ( Fig. 71 ). Hind femur with long hairs ( Fig. 69 ). Temple in dorsal view less rounded ( Fig. 67 ). Fore wing: proximal half of subhyaline, distal half brown fumous. ♀:? 8 mm M. hirtipes BRULLÉ, 1846

2 (1) Second submarginal cell less long, 3–SR 1.6 times as long as 2–SR ( Fig. 79). Hind femur with less long hairs ( Fig. 77 ). Temple in dorsal view receded ( Figs 75, 82 ). Fore wing evenly brown fumous. ♀: 8–8.5 mm M. lagopus ( KRIECHBAUMER, 1900)

The species is also near to M. rubriventre , for their distinction see key-couplets 8 (9)–9 (8).

Figures

Figs 74–83. Myosoma lagopus (KRIECHBAUMER), comb. n., female holotype: 74–81, female holotype of M. errotus SZÉPL.: 82–83: 74 = scape, pedicel and first flagellomere in outer-lateral view, 75 = head in dorsal view, 76 = head in lateral view, 77 = hind femur with indication of its hairs, 78 = claw, 79 = distal part of right fore wing, 80 = first discal cell of right fore wing, 81 = tergites 1–3, 82 = head in dorsal view, 83 = hind femur

Figs 66–73. Myosoma hirtipes BRULLÉ, female holotype: 66 = scape and pedicel in outer-lateral view, 67 = head in dorsal view, 68 = head in lateral view, 69 = hind femur and tibia with hairs, 70 = claw, 71 =distal part of left fore wing,72 =first discal cell of left fore wing, 73 = vein 1r–m of hind wing