Myosoma, BRULLE and Mysomatoides QUICKE. The, 1846

Papp, J., 2012, A Taxonomic Study Of The Myosoma Genus-Group With Description Of Amyosoma Cavei Sp. N. From Honduras (Hymenoptera: Braconidae: Braconinae: Braconini), Acta Zoologica Academiae Scientiarum Hungaricae 58 (1), pp. 1-29 : 6-9

publication ID	https://doi.org/ 10.5281/zenodo.5732026
DOI	https://doi.org/10.5281/zenodo.6908695
persistent identifier	https://treatment.plazi.org/id/03D0097D-E237-FF8F-E51E-DA5FDA7BFE2F
treatment provided by	Felipe
scientific name	Myosoma
status

Treatment

THE MYOSOMA View in CoL GENUS–GROUP

Within the tribe Braconini the Myosoma genus-group is distinguished by the following features: 1.) First tergite very long and narrow, two to five times as long as broad behind ( Fig. 83 View Figs 74–83 in QUICKE 1997: 172; Figs 29 View Figs 21–29 , 46 View Figs 38–47 , 56 View Figs 48–56 ) 2.) Second tergite considerably transverse, i.e. 3.3–4 times as broad behind as long medially, laterally narrowing ( Figs 37 View Figs 30–37 , 46 View Figs 38–47 , 65 View Figs 57–65 , 81 View Figs 74–83 , 91 View Figs 84–93 , 97 View Figs 94–97 ). 3.) Second submarginal cell long, i.e. 3–SR at least 1.5–1.6 times, frequently more than twice, longer than 2–SR ( Figs 26 View Figs 21–29 , 34 View Figs 30–37 , 42 View Figs 38–47 , 53 View Figs 48–56 , 62 View Figs 57–65 , 71 View Figs 66–73 , 79 View Figs 74–83 , 89 View Figs 84–93 , 95 View Figs 94–97 ).

The genus-group comprises three genera: Amyosoma VIERECK , Myosoma BRULLÉ and Mysomatoides QUICKE. The three genera and the species of Myosoma and Myosomatoides are separated by the features keyed:

1 (4) Second tergite with a distinct mid-longitudinal ridge ( Fig. 4 View Figs 1–13 in QUICKE 1994: 229). Hind femur and tibia flattened ( Fig. 3 View Figs 1–13 in QUICKE l.c.; Fig. 94 View Figs 94–97 ). The two Myosomatoides species are keyed after QUICKE (1994: 230) Myosomatoides QUICKE, 1994

2 (3) Wings uniformly brown. (= myersi QUICKE, 1994 syn. n., = pennipes MYERS, 1931 ) M. fasciatus ( BRULLÉ, 1846)

3 (2) Wings largely hyaline with apical third of the forewing brown ( Fig. 1 View Figs 1–13 in QUICKE l.c.) M. pennipes ( WESTWOOD, 1882)

4 (1) Second tergite without ridge. Hind femur and tibia not flattened ( Figs 24 View Figs 21–29 , 32 View Figs 30–37 , 40 View Figs 38–47 , 51 View Figs 48–56 , 60 View Figs 57–65 , 69 View Figs 66–73 , 77 View Figs 74–83 , 87 View Figs 84–93 ).

5 (6) Hind femur and tibia with short hairs as usually in Braconinae ( Fig. 32 View Figs 30–37 ). Vein 1r–m of hind wing weakly bent ( Figs 36 View Figs 30–37 , 45 View Figs 38–47 ) to almost straight ( Fig. 44 View Figs 38–47 ). See the key to the six species of the genus Amyosoma in VAN ACHTERBERG & POLASZEK (1996: 21). The seventh species, A. cavei sp. n., runs to A. rufescens ( QUICKE et INGRAM, 1993) with the help of this key, the distinction of these two species is presented at M. (A.) cavei sp. n. Amyosoma VIERECK, 1913

6 (5) Hind femur and hind tibia with long hairs ( Figs 24 View Figs 21–29 , 51 View Figs 48–56 , 69 View Figs 66–73 , 87 View Figs 84–93 ). Vein 1r–m of hind wing distinctly bent ( Figs 64 View Figs 57–65 , 73 View Figs 66–73 ) Myosoma BRULLÉ, 1846

7 (10) Fore wing: Second submarginal cell relatively short, i.e. 3–SR 1.5–1.6 times longer than 2–SR; SR1 1.5–1.7 times longer than 3–SR ( Figs 79 View Figs 74–83 , 89 View Figs 84–93 ). Temple in dorsal view strongly rounded to receded ( Figs 75, 82 View Figs 74–83 , 85 View Figs 84–93 ).

8 (9) First tergite twice longer than broad behind, scutum clearly broadening posteriorly ( Fig. 91 View Figs 84–93 ). Head in dorsal view relatively more transverse, 1.7 times as broad as long, temple strongly rounded ( Fig. 85 View Figs 84–93 ). Vein SR1 of fore wing approaching tip of wing ( Fig. 89 View Figs 84–93 ). ♀: 6 mm. – Brazil M. rubriventre BRULLÉ, 1846

9 (8) First tergite five times as long as broad behind, scutum less broadening posteriorly ( Fig. 81 View Figs 74–83 ). Head in dorsal view relatively less transverse, 1.6 times as broad as long, temple receded ( Figs 75, 82 View Figs 74–83 ). Vein SR1 of fore wing reaching tip of wing ( Fig. 79 View Figs 74–83 ). ♀: 8–8.5 mm. – Colombia M. lagopus ( KRIECHBAUMER, 1900)

10 (7) Fore wing: Second submarginal cell relatively long, i.e. 3–SR 1.8–2 times longer than 2–SR; SR1 1.1–1.3 times longer than 3–SR ( Figs 26 View Figs 21–29 , 53 View Figs 48–56 , 62 View Figs 57–65 , 71 View Figs 66–73 ). Temple in dorsal view rounded to less rounded ( Figs 22 View Figs 21–29 , 49 View Figs 48–56 , 58 View Figs 57–65 , 67 View Figs 66–73 ).

11 (14) Second submarginal cell very long, 3–SR 2.4–2.6 times as long as 2–SR ( Figs 26 View Figs 21–29 , 53 View Figs 48–56 ). Head in dorsal view a bit more transverse, 1.7 times as broad as long ( Figs 22 View Figs 21–29 , 49 View Figs 48–56 )

12 (13) Eye in dorsal view slightly less protruding and 1.5 times as long as temple ( Figs 38–39 View Figs 38–47 ). Hind femur less thick, 3.8 times as long as broad medially ( Fig. 40 View Figs 38–47 ). Metasoma black, second tergite laterally reddish. Fore wing brown fumous, distally (beyond pterostigma) subhyaline. ♀: 8 mm. – Bolivia M. brullei SZÉPLIGETI, 1906

13 (12) Eye in dorsal view slightly more protruding and 1.6 times as long as temple ( Fig. 49 View Figs 48–56 ). Hind femur thick. 2.9 times as long as broad medially ( Fig. 51 View Figs 48–56 ). Metasoma testaceous, tergites 6–8 black. Fore wing brown fumous, basally subhyaline. ♀: 10 mm. – Brazil M. errans ( SZÉPLIGETI, 1902)

14 (11) Second submarginal cell long, 3–SR 1.8–1.9 times as long 2–SR ( Figs 62 View Figs 57–65 , 71 View Figs 66–73 ). Head in dorsal view a bit less transverse ( Figs 58 View Figs 57–65 , 67 View Figs 66–73 ).

15 (16) Basal lobe of claw deep ( Fig. 70 View Figs 66–73 ). Eye in dorsal view 1.7 times longer than temple ( Fig. 67 View Figs 66–73 ). Scape in outer-lateral view longer than broad apically ( Fig. 66 View Figs 66–73 ). Metasoma reddish. ♀: 8.5 mm. – Ecuador M. hirtipes BRULLÉ, 1846

16 (15) Basal lobe of claw less deep ( Fig 61 View Figs 57–65 ). Eye in dorsal view 1.4 times longer than temple ( Fig. 58 View Figs 57–65 ). Scape in outer-lateral view cubic-form, i.e. as long as broad apically (cf. Fig. 48 View Figs 48–56 ). Metasoma testaceous, apically black. ♀: 9 mm. – Brazil, Belize M. fuscipenne BRULLÉ, 1846

Taxonomic remarks – 1.) The genera Amyosoma VIERECK, 1913 and Myosoma BRULLÉ, 1846 were considered as two valid taxa, however, recently QUICKE & INGRAM (1993: 317) and BELSHAW et al. (2001: 423) formally considered them congeneric. However, the genus Amyosoma is a valid genus accepting VAN ACHTERBERG & POLASZEK’ s (1996: 20) point of view on the distinction of the two genera in question.

2.) Ichneumon mutator FABRICIUS, 1775 was rearranged by BRULLÉ (1846: 453) into the genus Myosoma ; recently QUICKE & INGRAM designated it as the type species of their new genus Poecilobraconoides QUICKE et INGRAM, 1993. Furthermore, the species Iphiaulax bipartitus SZÉPLIGETI, 1905 is conspecific with P. mutator (FABRICIUS) ( QUICKE 1991: 175) distributed in Australia.

3.) The sixth species of the genus Myosoma , M. luteum , described by SZÉPLIGETI in 1913 was placed in the genus Cratocnema SZÉPLIGETI, 1914 ( QUICKE 1991: 184, PAPP 2000: 155). Concerning the new combination ( Cratocnema lutea ) QUICKE (l.c.) pointed out: “Very aberrant species of Cratocnema ...” As a result of the examination of the female lectotype of Myosoma luteum I confirm QUICKE’ s statement with the supplementary comment that it does not represent the genus Cratocnema but, perhaps, it belongs to an undescribed genus.

4.) The present taxonomic status of the Myosoma species by CAMERON see the checklist in Appendix.