Machimosauridae Jouve et al., 2016

Young, Mark T., Wilberg, Eric W., Johnson, Michela M., Herrera, Yanina, Brandalise, Marco de Andrade, Brignon, Arnaud, Sachs, Sven, Abel, Pascal, Foffa, Davide, Fernández, Marta S., Vignaud, Patrick, Cowgill, Thomas & Brusatte, Stephen L., 2024, The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha), Zoological Journal of the Linnean Society 200 (2), pp. 547-617 : 595

Machimosauridae Jouve et al., 2016 (Zoological Code)

Machimosauridae Johnson et al., 2020a: 117 (PhyloCode)

RegNum registration number 757.

Type genus: Machimosaurus von Meyer 1837 (emend. von Meyer 1838).

Etymology

‘The family of Machimosaurus ’. The stem Machimosaur - is based on the genus Machimosaurus . - idae, is a Latin patronymic suffix, introduced into zoological nomenclature, as -idae, by Kirby (1815: 88). Under Article 29.2 of the Zoological Code, the suffix - idae denotes a family rank within the family-group.

Geological range

Early Jurassic (early Toarcian) to Early Cretaceous (Hauterivian) ( Fanti et al. 2016, Johnson et al. 2020 a, Young and Sachs 2021). It is possible that the geological range of Machimosauridae spans from the Hettangian–Sinemurian to the late Barremian, if the newly described Early Jurassic specimen from Morocco is indeed a machimosaurid (as our phylogenetic analyses suggest) and if the incomplete late Barremian specimen from Colombia is also a machimosaurid ( Cortés et al. 2019, Hicham et al. 2023).

PhyloCode phylogenetic definition

Thelargestcladewithin Teleosauroidea containing Machimosaurus hugii ( von Meyer 1837) but not Teleosaurus cadomensis ( Lamouroux 1820) . Definition from Johnson et al. (2022b).

Reference phylogeny

Fig. 3 .

Zoological Code diagnosis and PhyloCode diagnostic apomorphies

Teleosauroid crocodylomorphs with the following unique combination of characters (5): between 50% and 65% of the premaxilla total length is posterior to the external nares (shared with Teleosaurinae ) (127.1); orbit longitudinal ellipsoidal in shape (reversal in Machimosaurini ) (shared with non-crocodylomorphs sampled; the early diverging crocodylomorph Carnufex ; the shartegosuchoid Sichuanosuchus ; the early diverging thalattosuchian Plagiophthalmosuchus ; the teleosauroids: the unnamed Chinese teleosauroid, Platysuchus , Seldsienean , and Aeolodon ; the metriorhynchoid clade Eoneustes + Metriorhynchidae ; the pholidosaurid Pholidosaurus purbeckensis ; and the hylaeochampsids Pietraroiasuchus and Iharkutosuchus ) (273.1); basisphenoid cultriform process exposed on the palate anterior to the quadrates (451.1); retroarticular process oriented posteriorly (reversal in Lemmysuchus and Yvridiosuchus ) (568.1); humerus proximal region very strongly deflected posteriorly and hooked (shared with Aeolodon ) (753.2).

Composition

The early diverging taxa Macrospondylus , Charitomenosuchus , and Deslongchampsina , and the subclade Machimosaurinae . The ‘Hettangian–Sinemurian’ taxon from Morocco forms a polytomy with the early diverging machimosaurids in our phylogenetic analyses.

Comments

Authorship: While nomen Machimosauridae was first used by Johnson et al. (2020a: 117), under the Zoological Code Jouve et al. 2016 is the nominal authority. The nominal author of a family-group is the author who first erected a family-group taxon that is valid (in fulfilment of Article 11), and in accordance with the Zoological Code Principle of Coordination applied to family-group names (Article 36.1). However, under the PhyloCode, the nominal authority is Johnson et al. (2020a).

Figures

Figure 3. The equal weights, no constraints strict consensus topology (C0). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S1 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/.