Philine sp. 2

Valdés, Ángel, 2008, Deep-sea “ cephalaspidean ” heterobranchs (Gastropoda) from the tropical southwest Pacific, Mémoires du Muséum national d'Histoire naturelle 196, pp. 587-792 : 725-726

Philine sp. 2

Figs 69C-E, 70

MATERIAL EXAMINED. — Indonesia. KARUBAR: stn DW 15 , Fiji. BORDAU 1 : stn DW 1408 , 16°02’S, 179°30’W, 550-561 m, 1 Kai Islands , 05°17’S, 132°41’E, 212-221 m, 1 dd. dd; stn DW 1410 , 16°06’S, 179°28’W, 400-410 m, 1 dd; stn DW New Caledonia proper. BATHUS 1 : stn DW 654 , Passe de Cana- 1440, 17°11’S, 178°43’W, 190-308 m, 1 dd; stn DW 1450 , 16°44’S, la, 21°17’S, 165°57’E, 237-298 m, 5 dd; stn DW 674 , Grande 179°58’W, 327-420 m, 1 dd; stn DW 1492 , 18°43’S, 178°23’W, Passe de Touho, 20°49’S, 165°19’E, 105-110 m, 1 dd; stn DW 430-450 m, 1 dd; stn DW 1494 , 18°55’S, 178°29’W, 240-319 m, 1 683, Passe de Hienghu, 20°35’S, 165°07’E, 380-400 m, 1 dd; dd; stn CP 1495, 18°53’S, 178°30’W, 420-445 m, 1 dd .

stn DE 700, Passe du Cap Baye, 20°57’S, 165°35’E, 160-222 Wallis and Futuna. MUSORSTOM 7: stn DW 585 , Wallis Ism , 2 dd . — BATHUS 2 : stn DW 733 , Passe de Kouaré , 22°55’S, land, 13°10’S, 176°13’W, 415-475 m, 1 dd; stn DW 601 , 13°19’S, 166°49’E, 520 m, 1 dd. 176°17’W, 350 m, 7 dd; stn DW 604 , 13°21’S, 176°08’W, 415- Norfolk Ridge. BERYX 11 : stn CP 21, 24°44’S, 168°07’E, 430- 420 m, 1 dd .

450 m, 8 dd; stn CH 41, 23°39’S, 168°00’E, 230-360 m, 1 dd. Tonga. BORDAU 2: stn DW 1508, Tongatapu, 21°02’S, 175°19’W, Loyalty Basin. BIOCAL: stn DW 77, 22°15’S, 167°15’E, 440 m, 555-581 m, 1 dd; stn DW 1509, 21°05’S, 175°22’W, 456-510 m, 3 1 dd. dd; stn DW 1540, SW of Tongatapu, 21°15’S, 175°14’W, 317-329 New Hebrides Arc. VOLSMAR: stn DW 51, Gemini Seamount, m, 1 dd; stn DW 1552, S of Nomuka Islands, 20°38’S, 174°58’W, 20°59’S, 170°03’E, 450 m, 2 dd. 491-500 m, 2 dd; stn DW 1614, seamount, 23°02’S, 175°51’W, Vanuatu. MUSORSTOM 8: stn DW 958, 20°21’S, 169°47’E, 429-549 m, 1 dd; stn DW 1617, 23°03’S, 175°53’W, 483-531 m, 497-570 m, 2 dd. 1 lv (Figs 69C-E).

DISTRIBUTION. — Collected from Fiji, Indonesia, New Caledonia, Tonga, Vanuatu and Wallis Island (Fig. 70), in 110-555 m, live in 483-531 m.

DESCRIPTION. — Shell morphology. Length 7 mm, width 4 mm (largest specimen examined). Shell internal, fragile, oval, with convex sides (Fig. 69C). The widest region of the shell is near the anterior end. Only 1 whorl visible, forming the entire shell. Anterior end of the shell rounded. Umbilicus closed. Aperture as long as the shell, wide anteriorly,

narrowing abruptly at about 5/6 of its length. Apex rounded, depressed, with a portion of the protoconch visible. Aperture lip conspicuously rising from the left side, forming a short wing posteriorly that rises above the apex. Columellar margin thickened. Sculpture of a number of punctuated spiral grooves. The punctuations are small, oval, and fused together with adjacent ones within each groove. The grooves are separated by gaps that are wider than the grooves. Occasionally narrower grooves are intercalated. Colour uniformly dirty white.

Anatomy. The digestive system contains 3 smooth gizzard plates, all small, simple and similar in shape and size (Fig. 69D). The radular formula is 12 x 2.1.1.1. 2 in a specimen from Tonga (BORDAU 2 stn DW 1617). The rachidian tooth is very small, rounded, with 3 denticles (Fig. 69E). The innermost lateral teeth are broad with a robust and curved cusp, bearing 8-10 strong denticles. The 2 outermost teeth are simple, with no denticles, elongate with a thin, curved cusp.

REMARKS. — Philine sp. 2 appears to be distinct from other species of the genus treated herein. However, all the specimens collected were either dried or devoid of soft parts, so anatomical examinations of the reproductive system were not possible. This species will remain unnamed until complete specimens become available for study.