Brachydesmus (Lophobrachydesmus) furcatus Lohmander, 1936,

Golovatch, Sergei, Evsyukov, Aleksandr & Reip, Hans, 2016, The millipede family Polydesmidae in the Caucasus (Diplopoda: Polydesmida), Zootaxa 4085 (1), pp. 1-51: 19-21

publication ID

http://doi.org/10.11646/zootaxa.4085.1.1

publication LSID

lsid:zoobank.org:pub:7F819439-FD77-4EF2-8E26-7B46F9B5FAAB

persistent identifier

http://treatment.plazi.org/id/03D23F44-F600-9D2F-FF07-5607FBE2C158

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scientific name

Brachydesmus (Lophobrachydesmus) furcatus Lohmander, 1936
status

 

Brachydesmus (Lophobrachydesmus) furcatus Lohmander, 1936 

Fig. 10View FIGURE 10, Map 3

Brachydesmus (Lophobrachydesmus) furcatus Lohmander, 1936: 16  (D).

Brachydesmus  ( Lophobrachydesmus  ) furcatus— Attems, 1940: 134 (D); Lang, 1959: 1795 (R). Brachydesmus furcatus  furcatus— Strasser, 1970: 204 (D); Lokšina and Golovatch, 1979: 384 (M); Talikadze, 1984: 143 (R). Brachydesmus (Lophobrachydesmus) furcatus exiguus Strasser, 1970: 204  (D), syn. n.

Brachydesmus furcatus  exiguus— Lokšina and Golovatch, 1979: 384 (M); Talikadze, 1984: 143 (R). Brachydesmus  furcatus— Chumachenko, 2012: 452 (R);

Material examined. 1 male, 5 juv. (ZMUM), RUSSIA, Krasnodar Prov., Gelendjik, Mikhailovskii Pass, Fagus  forest, 26.V.1956; 3 females (ZMUM), same locality and Pass, Fagus  forest, 5.VII.1956; 1 female, 3 juv. (ZMUM), same locality, Fagus  forest, under bark, 26.V.1956; 1 female, 2 juv. (ZMUM), Gelendjik, forest, 28.IV.1955; 1 female (ZMUM), Krasnodar Prov., Tuapse Distr., Georgievskoe Forestry, Castanea  forest, 7.V.1954; 1 female (ZMUM), same locality, SSW slope, Quercus  forest, 18.V.1954; 3 males, 6 juv. (ZMUM), Krasnodar Prov., Tuapse Distr., Cherepsinskoye Forestry, Quersus forest, S slope, 15.V.1954, all leg. M. Ghilarov, all det. H. Lohmander, det. C. Dziadosz, 1968; 1 female (ZMUM), Sochi, Khosta, Buxus  forest, 29.VI.1956, leg. M. Ghilarov; 2 females (ZMUM), same locality, Caucasian Nature Reserve, Taxus  and Buxus  forest, 15.V.1985, leg. S. Golovatch; 1 male, 1 female (ZMUM), same locality, Taxus  and Buxus  forest, III.2006, leg. Yu. Chumachenko; 1 female (ZMUM), Adygea, Guzeripl, Caucasian Nature Reserve, Pasture Abago S of Guzeripl, 1700 – 1850 m a.s.l., Abies  and Fagus  forest, Rhododendron  thicket, litter, 24 – 26.V.1985, leg. S. Golovatch; 1 male, 2 females (AE), Adygea, near Guzeripl, Abies  forest, 19.VI.2011, leg. D. Khisametdinova and E. Bortnikov; 1 female (ZMUM), Krasnodar Prov., Sochi, Krasnaya Polyana, 6 – 8.VI.1976; 1 female (ZMUM), same locality, 1900 m a.s.l., under stones, 29.VI.1956; 1 female (ZMUM), same locality, 600 m a.s.l., Castanea  forest, litter, 1.VI.1956, all leg. M. Ghilarov; 1 female (ZMUM), Krasnodar Prov., Sochi, Dagomys, 250 m a.s.l., Quercus, Corylus  , Fagus  etc. scrub, litter, logs, 18.V.1983; 2 females (ZMUM), Krasnodar Prov., Sochi, Caucasian Nature Reserve, Pslukh ca 20 km E of Krasnaya Polyana, Mt Aishkho-2, S slope, 1600 m a.s.l., Abies  , Fagus  , Betula  etc. forest, 12.VIII.1986; 1 male, 2 females (ZMUM), same locality, Mt Kogot, Abies  and Fagus  forest up to timberline, 1650 – 1850 m a.s.l., litter, under bark and stones, 18–20.V.1985; 1 female (ZMUM), Krasnodar Prov., Caucasian Nature Reserve, Krasnaya Polyana, 6 – 8.VI.1978, all leg. S. Golovatch; 2 females, 39 juv. (ZMUM), Krasnodar Prov., Sochi, Mt Bolshoy Akhun, Cave Akhunskaya, 19.VII.2015, leg. I. Turbanov; 9 juv. (ZMUM), Krasnodar Prov., Kabardinka, scrub, 2.VI.1956, leg. M. Ghilarov; 9 juv. (ZMUM), Krasnodar Prov., Severskaya Distr., 2 – 10 km S of Ubinskaya, 300 – 450 m a.s.l., Quercus  , Fagus  , Carpinus  etc. forest, litter and under bark, 3 – 4.VII.1986; 2 females, 22 juv. (ZMUM), same locality, Mt Derbiy ca 15 km SW of Ubinskaya, 800 – 850 m a.s.l., Quercus  , Fagus  , Fraxinus  etc. forest, litter and under bark, 2.VII.1986, all leg. S. Golovatch; 6 juv. (ZMUM), same locality, Ubinskoye Forestry,

Quercus  forest, 16.VI.1953, leg. M. Ghilarov; 2 males, 3 females (ZMUM), Krasnodar Prov., Goryachiy Klyuch, Cave Fanagoriyskaya, ca 12 km SW of Fanagoriyskoye, 30.X.1981; 1 male, 3 females, 3 juv. (ZMUM), same locality, Fagus  , Acer  , Picea  etc. forest, litter and under logs, 19.V.1983, all leg. S. Golovatch; 1 female (ZMUM), Krasnodar Prov., Goryachiy Klyuch, without data; 1 male (ZMUM), same locality, Kotkh Mt. Range, Quercus  forest, soil, 18.V.1956; 11 juv. (ZMUM), same locality, Pshaf Mt. Range, forest on limestone, 20.V.1956; 3 females (ZMUM), Pshaf Mt. Range, near “Belaya Latka”, Quercus  forest, 17.VI.1956; 3 females, 8 juv. (ZMUM), Pshaf Mt. Range, near “Belaya Latka”, S slope, Quercus  forest, 18.VI.1956; 8 females, 18 juv. (ZMUM), same locality, Kotkh Mt. Range, summit, S slope, Quercus  forest, 20.VI.1956; 2 females, 2 juv. (ZMUM), Kotkh Mt. Range, summit, S slope, Quercus  forest, 21.VI.1956; 1 male, 1 female (ZMUM), Kotkh Mt. Range, Quercus  forest, 18.VI.1956; 1 female (ZMUM), same locality, Canyon Bukovoe, 5.IV.1954; 7 juv. (ZMUM), Bolnichi, Mt Petushok, NE slope, Carpinus  forest, 4.VIII.1955; 1 juv. (ZMUM), Krasnodar Prov., Mt Ostraya, 1.VIII.1954; 1 female (ZMUM), same locality, 3.VIII.1954; 1 juv. (ZMUM), Kuban, Azovskaya, under Tkachovskoye, Quercus  forest belt, 24.VII.1955, all leg. M. Ghilarov, all det. H. Lohmander, det. C. Dziadosz, 1968.

2 males, 3 females (ZMUM), ABKHAZIA, Myussera Nature Reserve, 20 – 130 m a.s.l., Castanea  , Alnus  etc. forest, 8 – 10.IV.1983, leg. S. Golovatch; 1 male (SMNG 14685), Transcaucasus, Riza-See (= Lake Ritsa), 20.VI.1984, leg. W. Dunger.

Descriptive notes. Length ca 8.5 – 12 mm, width of midbody metazonae 0.8 – 1.3 mm (male, female), metazonite to prozonite width ratio <1.4 ( Fig. 10A – 10DView FIGURE 10). Live coloration usually uniformly pallid to light brown.

All characters as in B. assimilis  ( Fig. 10View FIGURE 10), except as follows. Paraterga poorly-developed, their fore margins and anterolateral corner largely broadly and regularly rounded, not angular; caudolateral corner mostly obtuseangled, pointed or nearly pointed and drawn behind rear tergal margin only in segments 16 – 18 ( Fig. 10A – 10DView FIGURE 10). Tergal setae mostly very short and subbacilliform. Metatergal sculpture especially superficial. Male prefemora not bulging laterad. Gonopod ( Fig. 10E – 10GView FIGURE 10) bipartite, strongly falcate, much as in B. pigmentatus  , but exomere (ex) with a conspicuous densely pilose strip, endomere (en) simple and bare, while tooth (t) at base of femorite very prominent and strongly rounded.

Remarks. The differences of B. furcatus exiguus Strasser (1970)  , described from a cave near Sochi, such as a slightly smaller and narrower body, a somewhat shorter endomere, a smaller pilose strip on the exomere and a less prominent tooth at the base of the gonofemorite ( Fig. 10 GView FIGURE 10) as compared to the sympatric nominate subspecies, seem to represent only individual variations. Hence we do not hesitate to formally synonymize B. furcatus exiguus  under B. furcatus  , syn. n. Moreover, the name B. f. exiguus Strasser, 1970  has long been jeopardized by the much older homonym, B. exiguus Brölemann, 1894  , from Italy ( Brölemann 1894).

Given the striking similarities observed in gonopod structure between B. (s. str.) pigmentatus  and B. (Lophobrachydesmus) furcatus  , the status of the latter subgenus is to be questioned.

Kingdom

Animalia

Phylum

Arthropoda

Class

Diplopoda

Order

Polydesmida

Family

Polydesmidae

Genus

Brachydesmus

Loc

Brachydesmus (Lophobrachydesmus) furcatus Lohmander, 1936

Golovatch, Sergei, Evsyukov, Aleksandr & Reip, Hans 2016
2016
Loc

Brachydesmus furcatus

Chumachenko 2012: 452
Talikadze 1984: 143
Loksina 1979: 384
1979
Loc

Brachydesmus

Talikadze 1984: 143
Loksina 1979: 384
Strasser 1970: 204
Strasser 1970: 204
Lang 1959: 1795
Attems 1940: 134
1940
Loc

Brachydesmus (Lophobrachydesmus) furcatus

Lohmander 1936: 16