Mniotype adusta adusta (Esper, 1790)

Volynkin, Anton V., Matov, Alexei Yu., Behounek, Gottfried & Han, Hui-Lin, 2014, A review of the Palaearctic Mniotype adusta (Esper, 1790) species-group with description of a new species and six new subspecies (Lepidoptera: Noctuidae), Zootaxa 3796 (1), pp. 1-32: 6

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Mniotype adusta adusta (Esper, 1790)


Mniotype adusta adusta (Esper, 1790) 

( Figs 15, 16View FIGURES 11 – 20, 59, 60View FIGURES 57 – 62, 88View FIGURES 87 – 96, 101View FIGURES 97 – 104)

Phalaena  Noctua adusta  Esper, [1790], Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen 4: 483 [1803], pl. 149 (Noct. 70), Figs 1–2View FIGURES 1 – 10 [1790] (Type locality: Germany, Frankfurt).

Synonymy: Noctua duplex Haworth, 1809  (Type locality: Great Britain); Noctua valida Hübner  , [1813] (Type locality: Europe); Hadena vultarina Freyer, 1832  (Type locality: unknown); Hadena chardinyi Duponchel  , [1838] 1836 (Type locality: [ France]); Hadena pavida Boisduval, 1840  (Type locality: Russia); Hadena baltica O. Hering, 1846  (Type locality: [ Poland] Danzig); Hadena adusta  var. grisescens Standfuss, 1893  (Type locality: Italy, Corsica Island); Crino aterrina Constantini, 1916  (Type locality: Italy); Hadena adusta  var. carpathica Kaucki, 1922  (Type locality: [ Poland] Malpolski); Crino adusta  subsp. lappona Rangnow, 1935 (Type locality: «Lappland»).

Material examined. 101 males, 78 females from Norway, Finland, Germany, Austria, Italy (Sicily), Ukraine (Poltava, Lvov areas), Russia (Karelia, Murmansk, Arkhangelsk, Saint-Petersburg, Jaroslavl, Tver, Moscow, Perm, Ekaterinburg, Orenburg, Krasnojarsk, Irkutsk, Chita and Kamchatka areas, Jakutia), E Kazakhstan (Saur and Tarbagatai Mts), W Mongolia (Bayan-Ölgyi and Hovd aimaks) (Colls ZISP, SZMN, AVB, GBG / ZSM).

Slides AV0516, AV0668, AV0682, AV0683, AV0713, AV0734, AV0739 Volynkin (males), AV0702, AV0721 Volynkin (females), glycerine preparations of both sexes by Volynkin and Matov.

Additional material examined. 1 male, Kazakhstan, Kurdai env., 2002, leg. Lind, slide 5838 Behounek (Coll. GBG / ZSM).

Note. The above specimen from north Tien Shan massif (SE Kazakhstan) has a habitus ( Fig. 20View FIGURES 11 – 20) and genitalia ( Fig. 66View FIGURES 63 – 68) close to those of the more northern populations. To clarify the status of the Tien Shan population, study of more extensive material is necessary.

Diagnosis. Wingspan 35–47 mm. The species is close to M. bathensis  and M. kobyakovi  . Externally M. adusta adusta  varies strongly in wing and body colouration. Ground colour of forewings varies from reddish brown to brown or grayish brown. M. adusta adusta  and M. bathensis  occur sympatrically over a large part of their ranges in North Europe, Ural and South Siberia. In some regions of South Siberia these two taxa occur sympatrically with a third species, M. kobyakovi  . All three taxa are highly variable externally; therefore, correct determination often requires study of the genitalia. In Europe, M. adusta adusta  usually has a less contrasting pattern and darker hindwings than European populations of M. bathensis  . In South Siberia, East Kazakhstan and West Mongolia, the unicolorous dark brown form of M. adusta adusta  is common. South Siberian populations of M. adusta adusta  were treated by Ronkay et al. (2001) as a subspecies ‘ M. adusta moesta (Staudinger, 1897)  ’; however, study of material from these areas showed that apart from a unicolorous dark form, a typical European form is common here. Thus, Siberian populations can not be treated as a distinct subspecies. The Siberian unicolorous specimens of M. adusta adusta  can be separated from M. bathensis  by their less contrasting pattern and darker reniform. Distinguishing external features of M. adusta adusta  from M. kobyakovi  are described in the diagnosis of the latter. The genitalia of M. adusta  ( Figs 59, 60View FIGURES 57 – 62, 88View FIGURES 87 – 96, 101View FIGURES 97 – 104) are close to M. kobyakovi  , M. juldussica  , M. adjuncta  and M. lama  . Distinguishing features from these taxa are described in their diagnoses.

Distribution. Eurasian. Europe, North and East Kazakhstan, European part of Russia, Ural, Siberia, north of Russian Far East, west Mongolia, north Tien Shan massif.


Zoological Institute, Russian Academy of Sciences


Siberian Zoological Museum


Goteburg Botanical Garden


Bavarian State Collection of Zoology