Mniotype dubiosa dubiosa (A. Bang-Haas, 1912)

Volynkin, Anton V., Matov, Alexei Yu., Behounek, Gottfried & Han, Hui-Lin, 2014, A review of the Palaearctic Mniotype adusta (Esper, 1790) species-group with description of a new species and six new subspecies (Lepidoptera: Noctuidae), Zootaxa 3796 (1), pp. 1-32: 14-15

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Mniotype dubiosa dubiosa (A. Bang-Haas, 1912)


Mniotype dubiosa dubiosa (A. Bang-Haas, 1912)  stat. n.

( Figs 43–45View FIGURES 41 – 50, 81, 82View FIGURES 81 – 86, 95View FIGURES 87 – 96, 117, 118View FIGURES 113 – 120)

Hadena lama  var. dubiosa A. Bang-Haas, 1912  , Deutsche Entomologische Zeitschrift Iris 26: 149 (Type locality: „Mustagata Yarkend“ [NW China, Xinjiang, Keng Tau Mts, Muztagh ata Mt.]).

Type material examined. Lectotype. Designated herein for Hadena lama  var. dubiosa  : male, “Mustagata Yarkend” (printed) / “ Hadena Lama Stgr.  v. Dubiosa  BH” (handwriten) / “B.H. 16 ” (handwriten) / “Preparation № MB. 355 Ch. Boursin” / pink label “Origin.” (printed) / (the lectotype deposited in coll NKMB). Paralectotype: female, „Mus-tag-ata Yarkend“ (printed) / “ Hadena lama dubiosa  ” (handwriten) / pink label “Origin.” (printed), slide Behounek 7659 f (deposited in coll NKMB).

Other material examined. 1 male, 5.vii. 2003, Kyrgyzstan, Sary-Dzhaz, floodplain of the river Kaingdy, near the mouth, V.V. Dubatolov, E.V. Nikolaeva [leg.] (Coll. SZMN); 1 male, 4.viii. 2003, Kyrgyzstan, S shore of Issyk- Kul lake, E Kara-Beldek Mts over the Teploklyuchenka, upper support point of the forestry, at light, V.V. Dubatolov, E.V. Nikolaeva [leg.] (Coll. SZMN); 2 males, 19.vii. 2003, Kyrgyzstan, Kashgaria, down stream of Archalsu river, V.V. Dubatolov & E.V. Nikolaeva leg. (Coll. SZMN); 1 male, [ Kyrgyzstan], 1906. vii. 5, Semirech. area, Naryn. Akulin [leg.] (Coll. ZISP); 1 male, [ Kyrgyzstan], Semirech. obl., Naryn, vi.–viii. 1905, unknown collector leg. (Coll. ZISP); 2 males, 4 females, [ Kyrgyzstan], Semirech. obl., Naryn, 10.v.- 1908, Akulin leg. (Coll. ZISP); 2 males, 4 females, 1909 / Ulug-Tuz, Sary-Kol Ridge, East Turkestan, Divnogorsky [leg.] (Coll. ZISP); 4 males, 4 females, China, Xinjiang, SW from Kashi, Keng Tau Mts, Outag loc., N 38 º 54.363 ', E 075º 13.788 '', h- 2650 m, Floriani leg., 4– 2013 (Coll. ASV); 3 males, 1 female, Kazakhstan, Prov. Almaty, Zailiskiy Alatau, 1700 m, 15 km S of Issyk, Issyk Lake, N 43 º 14 ', E077º 28 ', 30.v. 2006, leg. Balázs Benedek (Coll. BBT); 1 male, 07.vii. 2011, E Kazakhstan, Zaisan distr., Saur Mts, 24 km S Zaisan, upper stream of Bolshoi Zhemenei river, 1650 m, 47 ° 14 ’ N, 84 ° 56 ’ E. Titov S.V. leg. (Coll. AVB); 2 males, same locality, 17–19.vii. 2011, Yakovlev R.V. leg. (Coll. AVB); 3 males, 1 female, 18–19.vii. 2011, same locality, Titov S.V. leg. (Coll. AVB); 1 male, 2013, E Kazakhstan, East Kazakhstan area, Urdzhar district, Tarbagatai Ridge, 6,7 km N of Kyzymbet (Alekseevka) village, mesophilous shrubby slopes, 1300 m. 47 ° 18.365 ’ N, 81 ° 32.152 ’ E, Volynkin A.V., Titov S.V. & Černila M. leg. (Coll. AVB); 1 male, 15.viii. 2010, SE Kazakhstan, Zailijsky Alatau Ridge, middle stream of Assy river, 1700 m, Zhaman-Uj gorge, Egorov P.V. leg. (Coll. AVB); 1 female, 22–23.vii. 2009, SE Kazakhstan, Ketmen' Ridge, Amirsaj, Taranov B.T. leg. (Coll. AVB).

Slides AV0680, AV0685, AV0693, AV0694, AV0695, AV0704, AV0714, AV0715, AV0807, AV 1123 Volynkin (males), AV0716, AV0878 Volynkin, OP 2299 f, OP 2300 f Pekarsky (females).

Note. Previous authors have treated dubiosa  as a subspecies ( Hacker & Peks 1990; Ronkay et al. 2001) or a synonym (Hacker 1990) of M. lama  . Study of the type material demonstrates its specific status.

Diagnosis. Wingspan 37–40 mm. Head, thorax and abdomen brown. Ground colour of forewing brown. Outer margin of reniform dentate. Medial area shadowed around claviform and medial fascia. Terminal area dark brown. Hindwing pale brown. This species is externally close to M. juldussica  , but differs in the less dentate submarginal line, darker and more monotonous colouration of forewing and less expressed pattern. The correct determination of pale or damaged specimens often requires the study of the genitalia. The male genitalia of M. dubiosa  ( Figs 81, 82View FIGURES 81 – 86, 95View FIGURES 87 – 96) differ clearly from those of M. juldussica  in the narrower juxta with shorter serrated parts of its apical processes (in M. juldussica  distal half, but in M. dubiosa  only distal third serrate), narrower clavus strongly protruding from surface of sacculus, shorter ampulla, more rounded and apically elongated cucullus, obtuse and strongly dentate apical part of carina, shorter and less sclerotised eversible dorso-lateral bar of carina arising from the apical part of carina, and larger band of spinules on terminal diverticulum. The female genitalia ( Figs 117, 118View FIGURES 113 – 120) differ in the narrower ostium bursae, narrower and less curved ductus bursae, weakly sclerotised appendix bursae and posterior part of corpus bursae, larger conical anterior projection of anterior part of corpus bursae and smaller signum.

Distribution. Central Asian. East (Saur and Tarbagatai Mts) and South-East Kazakhstan, Tien Shan massif, Pamir and Alai Mts, nortwestern China (Xinjiang).


Siberian Zoological Museum


Zoological Institute, Russian Academy of Sciences