Scontromeryx falcidens ( Mazza et Rustioni, 2011 )

Van Der Geer, Alexandra A. E., 2014, Systematic revision of the family Hoplitomerycidae Leinders, 1984 (Artiodactyla: Cervoidea), with the description of a new genus and four new species, Zootaxa 3847 (1), pp. 1-32: 15-17

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Scontromeryx falcidens ( Mazza et Rustioni, 2011 )


Scontromeryx falcidens ( Mazza et Rustioni, 2011) 

Taxon A—Mazza & Rustioni 1999 (partim): p. 306 [SCT 70, SCT 103, 177].

Hoplitomeryx falcidens Mazza & Rustioni, 2011  — Mazza & Rustioni 2011 (partim): p. 1304, 1312, 1330, fig. 1, 2, 5, tables 1, 2 [SCT 70, SCT 82, SCT 103, SCT 177].

Holotype. Left hemimandible SCT 177 ( Figure 1View FIGURE 1, D, E, F, in Mazza & Rustioni 2011). Paratypes. Maxillary SCT 70, mandible fragments SCT 103, SCT 82.

Revised diagnosis. Small-sized, mesodont species. Lingual cusps tightly spaced in M 2 and M 3; metacone sometimes with small pillars on labial wall of upper molars; entostyle absent; mandible slender, with moderately sinuous horizontal ramus and convex ventral profile under cheek teeth corpus; paraconid absent or present only in unworn lower premolars; in p 2 protoconid and metaconid of same height; p 4 compressed mesiodistally, with enlarged and fairly complicated mesial trigonid and relatively small talonid; in lower molars labial cuspids triangular, widely spaced and somewhat backward-verging; metastylids and postentocristids well developed and protruding lingually in m 1; postentocristid hardly developed in m 3; ectostylid low and robust in m 1, low and blunt in m 3; hypoconulid and entoconulid with distal margins separated to halfway along the crown height; enamel smooth; weak cingulum at the base of the protoconid in m 1.

Differential diagnosis. Differs from Scontromeryx minutus  by having a mediodistally compressed lower p 4 (L/W ratio 1.3 vs 1.5), lacking a paraconid in lower premolars, and having smooth enamel on labial side of cheek teeth.

Derivation of name. According to Mazza & Rustioni (2011), the species is characterized by sabre-like upper canines (falx, gen. falcis, Latin, sickle, scythe; dens, Latin, masculine, tooth). However, maxillary SCT 177 lacks canines and the presence of a canine was based on material from the Gargano.

Preservation and deposition. Soprintendenza Archeologica dell’Abruzzo (Chieti, central Italy).

Type locality and horizon. Tortonian Scontrone Member of the Lithothamnium Limestone ( Patacca et al. 2008; 41 ° 45 ' 15.54 ''N, 14 °02' 13.14 ''E), outskirts of Scontrone, southern border of the National Park of Abruzzi, L’Aquila, central Italy.

Description. See Mazza & Rustioni (2011). Additional characters shown by paratypes revised after Mazza & Rustioni (2011). SCT 70 (M 2-3) is a maxillary fragment, SCT 82 (m 3) and SCT 103 (p 2 –m 1) are mandible fragments. m 3. The two mesial cuspids are not preserved (shown by SCT 82). Hypoconid markedly triangular and very inclined backwards. Entoconid fairly swollen at base of crown but postentocristid barely developed. Low, blunt ectostylid. Hypoconulid and entoconulid both tear-shaped and about the same size. Distal margins of these two conulids keep separated until about halfway the height of the crown. M 2–3. Small enamel pillars at the base of the metacone. Styles and ribs very strong. Metaconule with well-develop spur. Lingual cusps in tight contact with one another. No entostyle. M 3 with large metaconule.

Measurements. See Table 2.

TABLE 2. Dental measurements (L = length, W = width; in mm) of the mandibular (above) and maxillar fragments (below) from Scontrone, mandibular premolar / molar ratio and body mass estimation (BM in kg) based on lower molar row length after Damuth (1990) for selenodont browsers. * alveolar measurements. The specimens are sorted according to increasing size. Dental measurements from Mazza & Rustioni (2011, 1999, 1996).

  0.54* 18.2
S. apruthiensis   29.2
S. apruthiensis
S. apruthiensis   32.3
S. apruthiensis   33.3
S. apruthiensis   35.6
S. apruthiensis   37.6
S. apruthiensis
SCT58+SC T67
S. apruthiensis
  9.3 / 11.5














Scontromeryx falcidens ( Mazza et Rustioni, 2011 )

Van Der Geer, Alexandra A. E. 2014


Hoplitomeryx falcidens

Mazza & Rustioni 2011