Scontromeryx mazzai , Van Der Geer, Alexandra A. E., 2014

Van Der Geer, Alexandra A. E., 2014, Systematic revision of the family Hoplitomerycidae Leinders, 1984 (Artiodactyla: Cervoidea), with the description of a new genus and four new species, Zootaxa 3847 (1), pp. 1-32: 18

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Scontromeryx mazzai

sp. nov.

Scontromeryx mazzai  sp. nov.

Fig. 6View FIGURE 6 H in Mazza & Rustioni (2011)

Hoplitomeryx  sp. Leinders, 1984 – Rustioni et al. 1992: fig. 3 [uncatalogued skull fragment]; Mazza & Rustioni 1996: fig. 1 [SCT 17, redrawn after Rustioni et al., 1992: fig. 1].

Taxon C—Mazza & Rustioni 1999 (partim): p. 307 [ STCAbout STC 72, STCAbout STC 83].

Hoplitomeryx Leinders, 1984  — Mazza & Rustioni 1999 (partim): p. 305 [SCT 80].

Hoplitomeryx matthei Leinders, 1984  — Mazza & Rustioni 2011 (partim): p. 1320, 1321, 1330, fig. 6, tables 1, 2 [SCT 17, SCT 72, SC T 80, SCT 83].

Holotype. Skull fragment SCT 17.

Paratypes. Mandible fragments SC T 80, SCT 83, SCT 72.

Diagnosis. A hoplitomerycid with two pairs of orbital horns. Mandible markedly pachyostotic with brachyodont dentition. The lower m 2 and m 3 lack ectostylids, a cingulum and cementum and have rugose enamel.

Differential diagnosis. Differs from S. minutus  , S. falcidens  and S. apulicus  by pachyostosis of mandibular ramus and presence of orbital horns, differs from S. falcidens  and S. apulicus  by rugose enamel on its molars, differs from S. falcidens  by absence of ectostylid in m 2 and m 3 and by having brachyodont teeth. Differs from S. minutus  by absence of a cingulum and by rugose lingual enamel wall.

Derivation of name. Named in honour of Paul Mazza for his research on the artiodactyls from Scontrone.

Preservation and deposition. Soprintendenza Archeologica dell’Abruzzo (Chieti, central Italy).

Type locality and horizon. Tortonian Scontrone Member of the Lithothamnium Limestone ( Patacca et al., 2008; 41 ° 45 ' 15.54 ''N, 14 °02' 13.14 ''E), outskirts of Scontrone, southern border of the National Park of Abruzzi, L’Aquila, central Italy.

Description. See Mazza & Rustioni (2011). Additional characters shown by paratypes revised after Mazza & Rustioni (1999). The paratypes SCT 80 (m 3), SCT 83 (incomplete m 2, badly preserved m 3) and SCT 72 (incomplete m 2, m 3, alveoles p 2 –m 1) are fragmentary mandibles. The description here is after Mazza & Rustioni (1999) for SCT 72 and SCT 83. Horizontal ramus with marked pachyostosis. m 2 -m 3. Brachyodont without ectostylids. The ribs and the styles are slightly prominent on the lingual enamel walls. The rear wings of the protoconids are fused to the anterior wings of the entoconids. The labial crescents are obliquely arranged backwards and have a V-shaped outline in occlusal view. The posterior lobe is bicuspid with the labial lobe exceeding the lingual lobe in size. No cingula, no cementum. The enamel is rugose.

Measurements. See Table 2.

Remarks. Mazza & Rustioni (2011) attributed skull fragment SCT 17 and three mandible fragments (SCT 80, SCT 83, SCT 72) from Scontrone to Hoplitomeryx matthei  . However, these specimens cannot be attributed to this species. According to Mazza & Rustioni (2011) there is no evidence of the possible presence of the nasal horn core in skull fragment SCT 17, yet a nasal horn core is diagnostic for Hoplitomeryx matthei ( Leinders 1984)  . Furthermore, the orbital horn cores of SCT 17 show a combination of characters of Leinders’ (1984) horn core types I and II and constitutes thus a new horn core type, unlike any of Leinders’ (1984) types. As far as the mandibles are concerned, there is no evidence for the loss of the second premolar. This means that the presence of Hoplitomeryx matthei  cannot be confirmed among the material from Scontrone. In order to avoid confusion, the four Scontrone specimens which are excluded here from the H. matthei  material are attributed here to a new species as they cannot be attributed to any of the other Scontrone species defined by Mazza and Rustioni (2011).


Sichuan Teacher's College














Scontromeryx mazzai

Van Der Geer, Alexandra A. E. 2014



Leinders 1984


Hoplitomeryx matthei

Leinders 1984