PROBEEBEIDAE Boone, 1926

Fraaije, René H. B., Van Bakel, Barry W. M., Jagt, John W. M., Charbonnier, Sylvain, Schweigert, Guenter, Garcia, Géraldine & Valentin, Xavier, 2022, The evolution of hermit crabs (Crustacea, Decapoda, Anomura, Paguroidea) on the basis of carapace morphology: a state-of-the-art-report, Geodiversitas 44 (1), pp. 1-16 : 11

Family PROBEEBEIDAE Boone, 1926 ,

emended herein

Probeebeidae Boone, 1926b: 73 .

TYPE GENUS. — Probeebei (Boone, 1926) by monotypy.

OTHER GENERA INCLUDED. — Labidochirus Benedict, 1892 , Tisea Morgan & Forest, 1991 and Tylaspis Henderson, 1885 .

DIAGNOSIS. — Shield (excluding rostrum) strongly convex, well calcified, width equal to length or width exceeding length, with distinct bulges (i.e., keraial and massetic regions) laterally, strong to weaker spinose ornament. Rostrum well developed, exceeding lateral projections. Cervical and branchial grooves subparallel, encompassing small branchial regions. Posterolateral margins spinose. Posterior carapace well calcified, broadly inflated, with dense spinose ornamentation. Well-developed cardiac grooves encompassing cardiac region.

REMARKS. — The bulges on the shield of extant probeebeids ( Fig. 10 ) are quite similar to those of the most basal paguropsid, Eopaguropsis nidiaquilae (see Fraaije et al. 2012c: figs 2c; 3), which also has a spinose ornamentation on the gastric and lateral parts of the shield.

Figures

FIG. 10. — Transition from the Gastrodoridae to the Probeebeidae,featuring branchial condensation,reduction of rostrum,reduction of ornament on shield,inflation of posterior carapace,reduction of delineation of cardiac and reduction of midline carapace crest:A, Gastrodorus; B, Probeebei; C, Labidochirus; D, Tylaspis; E, Tisea.