Epipleoneura westfalli Machado, 1986

Pessacq, Pablo, 2014, Synopsis of Epipleoneura (Zygoptera, Coenagrionidae, “ Protoneuridae ”), with emphasis on its Brazilian species, Zootaxa 3872 (3), pp. 201-234 : 231-232

Epipleoneura westfalli Machado, 1986

( Figs. 16 , 43 , 58 )

Epipleoneura westfalli Machado 1986: 113 –118, figs. 1–3 (new species, description of male and female, illustration of male S 10 in lateral and posterior view, female prothorax in dorsal view, type material at ABMM). Lencioni 2005: 185, figs. 115 a–c (references, type locality, distribution, described stages, reproduction of illustrations of male S 10 in lateral and posterior view and female prothorax in dorsal view). Garrison et al. 2010: 352, 354, fig. 2338 (included in list of species, illustrations of epiproct in posterior view). Pessacq et al. 2012: 5 (included in list of Brazilian Protoneuridae , new record for Maranhão State).

Specimens examined. 2 ♂, 1 ♀, Brazil, Maranhão State, route São Luiz-Teresina (Teresina: 05º05’31”S, 42º48’12”W), km 340, 29 vii 1972.

Diagnosis. The cercus, epiproct, and genital ligula of this species are similar to those of E. kaxuriana , E. machadoi and E. ottoi ; for descriptions and further comments see under those species. The epiproct is strikingly similar to the atypical epiproct of a few specimens of E. machadoi from Baixo Guandu River ( Fig. 67 e), from which it can be distinguished by the smaller inner-basal teeth in E. westfalli , developed as a big triangular teeth or lobe in E. machadoi . The epiproct with divergent and curved branches characteristic of this species is also present in E. humeralis , but the genital ligula of these species is clearly different. The female posterior margin of pronotum ( Fig. 58 ) has an erect flat structure with a rounded lobe on each side and a deep U cleft between them, which further differentiates it from E. kaxuriana , E. machadoi , and E. ottoi , but is similar to the bilobed margin of pronotum of E. humeralis .

Distribution. Northern Brazil in Maranhão and Rondônia States (type locality, Ji Paraná, Rondônia State, about 10°52'51"S 61°56'31"W).

Notes. Some variation was observed in the epiproct of this species; as mentioned by Machado (1986), the branches are divergent and with apex acute or slightly divergent with stout apex.

Figures

FIGURES 16 – 18. Male caudal appendages, a: lateral view; b: posterior view, c: latero-posterior view. Figures 17 and 18 modified from Garrison et al. (2010) and De Marmels (1985) respectively. CT: cercus inner-basal tooth.

FIGURES 28 – 47. Genital ligula, lateral and ectal view. Figures 29, 33, 47 modified from Garrison et al. (2010). PLP: posterolateral projection.

FIGURES 53 – 66. Female pronotum in lateral, dorsal, antero-lateral, anterior postero-lateral and posterior view. Figures 57, 64 and 65 modified from Garrison et al. (2010), Rácenis (1960) and Machado (2005) respectively.