Homotropus klopfsteinae Johansson, 2020

Morishita, Shunsuke & Watanabe, Kyohei, 2025, Review of the genera Homotropus Förster and Syrphoctonus Förster (Hymenoptera, Ichneumonidae, Diplazontinae) from Japan, Zootaxa 5588 (1), pp. 49-76 : 62-64

Homotropus klopfsteinae Johansson, 2020

( Figs 8 A, B )

Homotropus klopfsteinae Johansson, 2020: 74 .

Specimens examined. JAPAN: [Hokkaido] 1M, Kamikawa Town, Sounkyo, Ginsendai 8–9 IX 2014, S. Shimizu leg. ( KPMNH) .

Diagnosis. Antenna with 23 (in female) or 25 (in male) flagellomeres. Tyloids absent in male. Face coriaceous and matt; sparsely punctate. Clypeus depressed apically; robes rounded at outer corners. Mesoscutum coriaceous and matt; densely punctate. Mesopleuron coriaceous and matt; densely punctate anteriorly and ventrally. Propodeum finely rugulose; lateral longitudinal and pleural carina present; petiolar area surrounded by carina ( Fig. 1 C ). Fore wing length 7.8 (in female) or 8.0 (in male) mm. Areolet present. Dorsal part of hind coxa coriaceous and matt. Metasoma dorsoventrally depressed in both sexes. Latero-median carina present on basal 0.8 of T I. T I 2.0 (in female) or 1.8 (in male) × as long as maximum width; strigose anteriorly; otherwise coriaceous. T II 1.3 (in female) or 1.0 (in male) × as long as maximum width. Face black with pair of yellow markings along each inner orbit ( Fig. 12 B ). Clypeus entirely yellow in both sexes ( Fig. 8 B ). Malar space entirely black in both sexes ( Fig. 8 B ). Yellow shoulder markings of mesoscutum present ( Fig. 8 A ). Scutellum entirely yellow. All coxae entirely black in females. Fore coxa black with yellow apex, mid and hind coxae entirely black in males ( Fig. 8 A ). Hind tibia yellow, darkened apically ( Fig. 8 A ). Metasoma entirely black in both sexes.

Description based on Japanese specimen. Male (n=1). Body covered with silverly setae; length 10.0 mm.

Head 0.51 × as long as wide in dorsal view; coriaceous; sparsely punctate. Clypeus 2.0 × as maximum width as high; depressed apically; robes rounded at outer corners. Face 2.3 × as maximum width as high; weakly convex medially in lateral view; separated from clypeus by shallow clypeal sulcus. Malar space 1.0 × as long as basal mandibular width.POL 1.55 × as OD. OOL 1.15 × as OD. POL 1.40 × as long as OOL.Antenna with 25 flagellomeres. Tyloids absent. FL I 1.2 × as long as FL II. Maxillary palp IV1.3 × as long as Maxillary palp V.

Mesosoma densely punctate. Lateral aspect of pronotum coarsely rugulose anteriorly and ventrally.Mesopleuron smooth below speculum. Sternaulus weakly impressed. Propodeum finely rugulose; lateral longitudinal and pleural carina present; petiolar area surrounded by carina ( Fig. 1 C ). Fore wing length 8.0 mm. Areolet present. Vein 1cu-a of fore wing postfurcal to vein M&RS. Hind wing with nervellus intercepted below middle. Hind wing with 9 distal hamuli. Dorsal part of hind coxa coriaceous and matt. Hind femur 6.25 × as long as maximum depth in lateral view. Hind tibia 8.3 × as long as maximum depth in lateral view. Ratio of length of hind first to fifth tarsomeres 1.0: 0.7: 0.4: 0.2: 0.3.

Metasoma coriaceous; dorsoventrally depressed. T I rectangular in dorsal view; 1.8 × as long as maximum width; strigose anteriorly; otherwise coriaceous; latero-median carina present on basal 0.8. T II 0.95 × as long as maximum width; strigose anteriorly.

Coloration ( Figs 8 A, B ). Body (excluding wings and legs) black. Face with pair of yellow markings along each inner orbits. Palpi, mandible except for apex, subtegular ridge, tegula, upper mesepisternum, shoulder markings of mesoscutum, and scutellum yellow. Legs black. Apex of fore coxa, trochanters, trochantelli, and base of hind tibia yellow. Fore and mid femora, tibiae, and tarsomeres yellowish-brown to brown. Base and apex of hind femur brown.

Distribution. Japan (Hokkaido). Outside Japan, this species is distributed in Sweden ( Johansson 2020).

Bionomics. In Japan, adults were collected in broad-leaved forest at altitudes of ca. 1,000 meters.

Remarks. This is the first record of the male of this species and the first record of this species from Japan.

Figures

FIGURE 1. Japanese Homotropus spp., females—A, B: T I, dorsal view; C: propodeum, dorsal view.A: H.formosus Klopfstein, 2014; B: H. sundevalli (Holmgren, 1858); C: H. klopfsteinae Johansson, 2020.

FIGURE 8. Homotropus klopfsteinae Johansson, 2020, male―A: lateral habitus; B: head, frontal view.

FIGURE 12. Homotropus signatus (Gravenhorst, 1829), female (A–D) and male (E, F)―A, E: lateral habitus; B, F: head, frontal view; C: scutellum, dorsal view; D: T II to apex of metasoma, dorsal view.