Notocupes, Ponomarenko, 1964

Strelnikova, O. D. & Yan, E. V., 2023, ON SPLITTING OF THE GENUS NOTOCUPES (COLEOPTERA: ARCHOSTEMATA): NEW DATA ON MORPHOLOGY AND TAXONOMY, Far Eastern Entomologist 488, pp. 1-28 : 19-20

Key to species of the genus Notocupes

( N. baojiatunensis , N. kezuoensis , N. laetus , N. laiyangensis , N. lini , N. ludongensis , N. neli , N. ohmkuhnlei , N. reticulatus , N. shiluoensis , N. tuanwangensis and N. validus does not include in a key due to lack of observable diagnostic characters in published sources).

1. Sides of pronotum and/or elytron are dentate at basal third of their length ........................ 2

– Pronotum and elytron without denticles ............................................................................. 11

2. Abdominal sternites have depressions along their anterior margins and paired triangular lateral depressions. Body covered with two types of tubercles: small and large. Abdomen weakly narrowing basally (except in N. latus , which has only small tubercles and has abdomen not narrowing basally) ....................................................................................... 3

– There are no paired triangular depressions on abdominal sternites. Body covered with uniform tubercles. Abdomen not narrowing basally ......................................................... 8

3. Body covered with single type small (0.01–0.02 mm in diameter) tubercles, abdomen not narrowing basally ................................................................................................... N. latus

– Body has two types of tubercles: small (0.01–0.02 mm in diameter) and large (0.04–0.06 mm in diameter), abdomen slightly narrowing basally ..................................................... 4

4. Pronotal width to length ratio more than 2; metaventrite`s width is equal to its length ....... ...……………………………………………………………………….. N. khasurtyiensis

– Pronotal width to length ratio less than 2; metaventrite`s width to length ratio 1.5–1.8 ...... 5

5. Epipleuron without cells, more than 10 times narrower than elytra ...……..…. N. excellens

– Epipleuron with cells, less than 10 times narrower than elytra ....…………….…………... 6

6. Epipleuron strongly narrows at basal third, 4 times narrower than its average width ............ ...……………………………………………………………………….….. N. mongolicus

– Epipleuron gradually narrows at basal third, 1.3 times narrower than its average width ..... 7

7. Epipleuron narrow, 8.5 times narrower than elytron, the apical abdominal sternite to penultimate one length ratio is 3.1 ................................................................... N. caudatus

– Epipleuron wide, 5.9 times narrower than elytron, the apical abdominal sternite to penultimate one length ratio is 2.4 ........................................................ N. daohugouensis

8. Antennal bases shifted to dorsal side ( Fig. 38 ) .................................................... N. pulcher

– Antennae attached laterally in front of eyes .....………………………………………….... 9

9. Pronotum and basal half of elytron`s epipleural margin with distinct denticles. One elytral cell has about 2–3 maculae ......................................................................... N. cyclodontus

– Basal half of elytron`s epipleural margin not dentate; pronotum laterally dentate. One elytral cell has no less than 3 maculae.…………………………………….…………... 10

10. Pronotum laterally slightly dentate, one elytral cell has 3–4 maculae ................... N. rudis

– Pronotum laterally strongly dentate, оne elytral cell has 5–8 maculae ............... N. robustus

11. Head rounded; anterior margin of prosternum have two denticles .................... N. alienus

– Head rectangular, not narrowing anteriorly or posteriorly; prosternum without denticles.… .......................................................................................................................................... 12

12. Pronotum wide, width to length ratio 1.8–2; 25–30 cells in one elytral row ................... 13

– Pronotum less wide, width to length ratio 1.2–1.3; 20–25 cells in one elytral row.…....... 14

13. Large beetles, body length more than 20 mm. Elytral cells medium-sized, about 25 cells in one elytral row. Epipleuron wide, strongly and abruptly narrowing at basal third. Metaventrite`s width 1.8 times exceeds length. Length of apical abdominal sternite two times as long as wide ................................................................................. N. diazromerali

– Beetle less than 10 mm in length. Elytral cells small, about 30 cells in one elytral row. Epipleuron narrow, gradually narrowing at basal half. Metaventrite`s width and length are equal. Width of apical abdominal sternite 1.4 less its length ........................... N. pingi

14. Apical abdominal sternite to penultimate one length ratio more than three ( Fig. 60 ) ........ ……………………………………..…………………………………….….. N. lapidarius

– Apical abdominal sternite to penultimate one length ratio 2.3–2.8 .................................... 15

15. Cuticular tubercles not larger than 0.03 mm, density no less than 760 tb/mm 2. Epipleuron narrow, weakly and gradually narrowing at elytron basal third ( Fig. 7 ) ........ …………………………………………………………………………….… N. picturatus

– Larger tubercles: 0.03–0.05 mm in diameter, density no more than 320 tb/mm 2. Epipleuron of medium width, abruptly narrowing at elytron basal third ( Figs 27, 28 ) ........ N. elegans

Figures

Figs 38–41. Antennae morphology. 38 – filiform of Notocupes pulcher; 39 – moniliform of Notocupes excellens; 40 – weakly serrated of Rhabdocupes minisculus; 41 – filiform of Brachilatus nigrimonticola. Scale bar = 1 mm.

Figs 57–60. Abdomen morphology. 57 – Rhabdocupes rostratus; 58 – Conexicoxa longicollis; 59 – Brachilatus caducus; 60 – Notocupes lapidarius. Scale bar = 1 mm.

Figs 5–8. Representatives of the four investigated genera. 5 – Rhabdocupes protensus (Tan et al., 2006); 6 – Conexicoxa epicharis (Tan, Ren et Liu, 2005); 7 – Notocupes picturatus Ponomarenko, 1964; 8 – Brachilatus longicoxa (Soriano et Martinez-Delclòs, 2006). Scale bar = 1 mm.

Figs 25–28. Elytron venation types of five investigated genera of Archostemata. 25, 26 – Zygadenia alexrasnitsyni Strelnikova et Yan, 2021: 25 – photograph; 26 – interpretative linedrawing; 27, 28 – Notocupes elegans Ponomarenko, 1994: 27 – photograph; 28 – interpretative linedrawing. Elytral fields marked with Roman numerals, veins – with Arabic. Scale bar = 1 mm.