Megascolecidae

Tsai, Chu-Fa, Shen, Huei-Ping, Tsai, Su-Chen, Lin, Kang-Jieh, Hsieh, Hwey-Lian & Yo, Shao-Pin, 2009, A checklist of oligochaetes (Annelida) from Taiwan and its adjacent islands, Zootaxa 2133, pp. 33-48: 38-45

publication ID

http://doi.org/ 10.5281/zenodo.188447

persistent identifier

http://treatment.plazi.org/id/03D6CB65-FFCE-3676-25DF-FE8037266B6C

treatment provided by

Plazi

scientific name

Megascolecidae
status

 

Family Megascolecidae 

36. Pontodrilus litoralis  (Grube, 1855)

Pontodrilus litoralis  - Shen et al., 2005 a: 12. Beaches of Sanshui and Chitou, Penghu Island, Haomeiliao, Chiayi County, and Chiku Lagoon, Tainan County; - James et al., 2005: 1022. Marine shoreline near coral reefs, Wanlitong, Pingtung County and mangroves at Kaoping River Estuary, Linyuan, Kaohsiung County; - Blakemore et al., 2006: 230.

37. Pithemera bicincta  (Perrier, 1875)

Pheretima bicincta  - Gates, 1959: 4. Yangmingshan, Taipei City.

Pithemera bicincta  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 288; - Shen & Tsai, 2002 b: 3. Lanyu Island; - Chen et al., 2003: 58. Nanao and Fushan Botanical Garden, Ilan County; - Blakemore et al., 2006: 230.

38. * Pithemera lanyuensis  Shen & Tsai, 2002

Pithemera lanyuensis  - Shen & Tsai, 2002 a: 2 and 2002 b: 3. Lanyu Island; - Chang & Chen, 2004: 219; - Blakemore et al., 2006: 230.

39. Perionyx excavatus  Perrier, 1872

Perionyx excavatus  - Kobayashi, 1938: 201 and 1939 b: 779. Hsinchu; - Gates, 1959: 19. Suao, Ilan County; - Kuo, 1995: 189. Chasili village, Chungli, Taoyuan County; - Shih et al., 1999: 436; - Tsai et al., 2000 a: 286; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chuang et al., 2002: 68. Taipei County; - Tsai et al., 2004 a: 6. Coastal plain to the Central Mountain Range to the elevation of 1100 m in central Taiwan; - Blakemore et al., 2006: 230.

40. Amynthas aspergillum  ( Perrier, 1872)

Perichaeta takatorii Goto & Hatai, 1898: 76  . Taipei City.

Pheretima (Pheretima) aspergillum  - Michaelsen, 1929: 158. Formosa.

Pheretima takatorii  - Kobayashi, 1939 a: 659, 1939b: 778, 1940 a: 120, 1940 b: 274, and 1940 c: 391. Taipei and Hsinchu.

Pheretima aspergillum  - Gates, 1959: 1. Chaochou, Pingtung County; - Tsai, 1964: 29. Sand bank of Danshui River and National Taiwan University campus, Taipei.

Amynthas aspergillum  - Shih et al., 1999: 436; Chuang et al., 2002: 68. Taipei City, Hsinchu and Taipei counties; - Blakemore et al., 2006: 227.

Amynthas aspergillus  - Tsai et al., 2000 a: 286; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Shen & Tsai, 2002 b: 4. Lanyu Island; - Chen et al., 2003: 58. Jiaohsi, Nanao and Suao, Ilan County; - Tsai et al., 2004 a: 5. Coastal plain to the elevation of 600 m in central Taiwan.

41. * Amynthas bilineatus  Tsai & Shen, 2007

Amynthas bilineatus Tsai et al., 2007: 366  . Hill slope along Dongyen Creek, Nantou County.

42. * Amynthas binoculatus  Tsai, Shen & Tsai, 1999

Amynthas binoculatus Tsai et al., 1999: 41  . Wufong, Taichung County; - Tsai et al., 2000 a: 286; - Tsai et al., 2004 a: 6. Lower peripheral hills at an elevation of 300 m; - Blakemore et al., 2006: 228.

43. * Amynthas candidus  ( Goto & Hatai, 1898)

Perichaeta candida Goto & Hatai, 1898: 77  . Taipei City.

Pheretima candida  - Gates, 1959: 18.

Amynthas candidus  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 286; - Blakemore et al., 2006: 228.

44. Amynthas carnosus  (Goto & Hatai, 1899)

Amynthas carnosus  - Shen et al., 2005 b: 97. Tunghou, Hsiaoi Village, Wulai, Taipei County, and Hsiaojiaohsi and Rt. 33 - 1 near Meihua Lake, Ilan County; - Blakemore et al., 2006: 228.

45. * Amynthas catenus  Tsai, Shen & Tsai, 2001

Amynthas catenus Tsai et al., 2001: 279  . Mt. Hohuan between Hualien and Nantou counties; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 228.

46. * Amynthas chilanensis  Tsai & Tsai, 2007

Amynthas chilanensis Tsai et al., 2007: 362  . Chilan, Ilan County.

47. Amynthas corticis  (Kinberg, 1867)

Pheretima diffringens (Baird, 1869)  - Kobayashi, 1939 a: 660, 1939b: 778, 1940 a: 120, 1940 b: 274, and 1940 c: 390. Hsinchu; - Gates, 1959: 6. Taipei; - Tsai, 1964: 2. Sand bank of Danshui River and National Taiwan University campus, Taipei; - Chen & Shih, 1996: 54. Fushan Botanical Garden, Ilan County.

Amynthas corticus  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 286; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chen et al., 2002: 77. Fushan Botanical Garden, Ilan County; - Chuang et al., 2002: 68. Taipei, Taoyuan, Hsinchu, and Ilan counties; - Shen & Tsai, 2002 b: 3. Lanyu Island; - Chen et al., 2003: 58. Sanhsing, Tungshan, Yuanshan, Jiaohsi, Nanao, Suao, Fushan Botanical Garden, Datung, and Taipinshan, Ilan County.

Amynthas cortices  - Chuang et al., 2002: 70. Taipei, Taoyuan, Hsinchu, and Ilan counties; - Chen et al., 2004: 118. Six northern counties.

Amynthas corticis  - Tsai et al., 2004 a: 6. Coastal plain to the Central Mountain Range to the elevation of 2100 m in central Taiwan; - James et al., 2005: 1023. Mt. Nanren, Jioupeng, and Nanhe, Chunrih, Pingtung County; - Blakemore et al., 2006: 228.

48. * Amynthas cruxus  Tsai & Shen, 2007

Amynthas cruxus Tsai et al., 2007: 368  . Tengchih, Shihshan Forest Road and Chuyunshan Nature Reserve, Kaohsiung County.

49. * Amynthas exiguus aquilonius  Tsai, Shen & Tsai, 2001

Amynthas exiguus aquilonius Tsai et al., 2001: 277  . Mt. Hohuan, Hualien County and Rueyen Creek Nature Reserve, Nantou County; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 228.

50. * Amynthas fenestrus  Shen, Tsai & Tsai, 2003

Amynthas fenestrus Shen et al., 2003 b: 487  . Rueyen Creek Nature Reserve and Mt. Beidongyen, Nantou County; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 228.

51. Amynthas gracilis  (Kinberg, 1867)

Pheretima  sp. Takahashi, 1931: 10 and 1932: 344. Taipei City. (According to the structure described, it was later identified as Pheretima hawayana  ).

Pheretima hawayana (Rosa, 1891)  - Kobayashi, 1939 b: 778, 1940a: 120, 1940 b: 274, and 1940 c: 390. Hsinchu; - Tsai, 1964: 9. Sand bank of Danshui River, National Taiwan University campus, Yingko, and Wanhua, Taipei; - Kuo, 1995: 184. Hsinpu, Hsinchu; - Chen & Shih, 1996: 54. Fushan Botanical Garden, Ilan County.

Amynthas gracilis  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 286; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chen et al., 2002: 77. Fushan Botanical Garden, Ilan County; - Chuang et al., 2002: 68. Taipei, Taoyuan and Ilan counties; - Shen & Tsai, 2002 b: 3. Lanyu Island; - Chen et al., 2003: 58. Sanhsing, Tungshan, Yuanshan, Jiaohsi, Nanao, Suao, Datong, and Gueishan (Turtle) Island, Ilan County; - Chen et al., 2004: 118. Six northern counties; - Tsai et al., 2004 a: 6. Coastal plain to the Central Mountain Range to the elevation of 2100 m in central Taiwan; - James et al., 2005: 1024. Caopu and Gaoshih, Pingtung County; - Blakemore et al., 2006: 228.

52. * Amynthas hohuanmontis  Tsai, Shen & Tsai, 2002

Amynthas hohuanmontis Tsai et al., 2002: 758  . Mt. Hohuan, Nantou County; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 228.

Remarks: A. hohuanmontis  was originally described as an athecate earthworm (Tsai et al. 2002). However, examination of specimens from successive collections showed that this species has spermathecae from three pairs in VI-VIII to absence, and the structure of the prostate glands varies from normal, asymmetrical, to absence, which also indicates parthenogenetic degeneration.

53. Amynthas hupeiensis  (Michaelsen, 1895)

Pheretima hupeiensis  - Kobayashi, 1939 a: 660, 1939b: 778, 1940 a: 120, 1940 b: 274, and 1940 c: 390.

Hsinchu; - Tsai, 1964: 11. Sand bank of Danshui River, Taipei City.

Amynthas hupeiensis  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 287; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chen et al., 2002: 77. Fushan Botanical Garden, Ilan County; - Chuang et al., 2002: 68. Taipei, Hsinchu and Ilan counties; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 228.

54. Amynthas incongruus  ( Chen, 1933)

Pheretima incongrua  - Gates, 1959: 13. Taipei; - Tsai, 1964: 19. National Taiwan University campus, Taipei City.

Amynthas incongruus  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 287; - Tsai et al., 2004 a: 6; - James et al., 2005: 1025. Mt. Nanren, Pingtung County; - Blakemore et al., 2006: 228.

Amynthas incongrua  - Chuang et al., 2002: 69. Taoyuan and Hsinchu counties; - Chen et al., 2003: 58. Yuanshan, Jiaohsi, Suao, Fushan Botanical Garden, Datong, and Gueishan (Turtle) Island, Ilan County.

55. * Amynthas lini  Chang, Lin, Chen, Chuang & Chen, 2007

Amynthas lini Chang et al., 2007 b: 234  . Taipei, Ilan, Taoyuan, Hsinchu, Nantou, and Chiayi counties.

56. * Amynthas meishanensis  Chang, Lin, Chen, Chuang & Chen, 2007

Amynthas meishanensis Chang et al., 2007 b: 236  . Meishan, Chiayi County.

57. Amynthas minimus  (Horst, 1893)

Pheretima zoysiae Chen, 1933  - Kobayashi, 1940 b: 274 and 1940 c: 391. Hsinchu; - Kobayashi, 1941: 378. Taiwan.

Amynthas minimus  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 287; - Blakemore et al., 2006: 228.

Remarks: A. minimus  was first collected by Kobayashi (1940 b, c). Recently, it was also collected by us from the No. 100 Forest Road (elevation 1060 m), Ilan County on 17 October 2002 and Hushan Temple (elevation 140 m), Douliou, Yunlin County on 12 June 2007.

58. Amynthas morrisi  (Beddard, 1892)

Pheretima morrisi  - Kobayashi, 1939 b: 778, 1940a: 120, 1940 b: 274, and 1940 c: 390. Hsinchu; - Tsai, 1964: 17. Yingko, Taipei County.

Amynthas morrisi  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 287; - Chuang et al., 2002: 69. Hsinchu County; - Tsai et al., 2004 a: 6. Coastal plain to the Central Mountain Range to the elevation of 1100 m in central Taiwan; - Blakemore et al., 2006: 228.

59. * Amynthas nanrenensis  James, Shih & Chang, 2005

Amynthas nanrenensis James et al., 2005: 1008  . Mt. Nanren, Pingtung County; - Blakemore et al., 2006: 228.

60. * Amynthas nanshanensis  Shen, Tsai & Tsai, 2003

Amynthas nanshanensis Shen et al., 2003 b: 482  . Mountain slop along Nanshan Creek and Mt. Beidongyen, Nantou County; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 228.

61. Amynthas papilio  (Gates, 1930)

Amynthas papilio  - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chuang et al., 2002: 68. Taipei and Hsinchu counties; - Chen & Chuang, 2003: 90. National Taiwan University and Hsinchu Girl High School campuses.

Amynthas papilio papilio  - Blakemore et al., 2006: 228.

62. Amynthas papulosus  ( Rosa, 1896)

Pheretima papulosa sauteri Michaelsen, 1922: 26  . Chiahsien (= Dorf Kosempo), Kaohsiung County; - Kobayashi, 1939 a: 660, 1939b: 778, 1940 a: 120, 1940 b: 274, and 1940 c: 390. Hsinchu.

Pheretima papulosa  - Gates, 1959: 18.

Amynthas papulosus  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 287; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chen et al., 2002: 77. Fushan Botanical Garden, Ilan County; - Chuang et al., 2002: 68. Taipei, Taoyuan, Hsinchu, and Ilan counties; - Chen et al., 2003: 58. Jiaohsi, Suao and Gueishan (Turtle) Island, Ilan County; - Tsai et al., 2004 a: 6. Tsaotun, Nantou County; - Blakemore et al., 2006: 228.

63. * Amynthas penpuensis  Shen, Tsai & Tsai, 2003

Amynthas penpuensis Shen et al., 2003 b: 481  . Mountain slops along Penpu and Nanshan creeks, Nantou County; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 228.

64. * Amynthas polyglandularis  ( Tsai, 1964)

Pheretima polyglandularis Tsai, 1964: 30  . Hill behind the 8 th dormitory of National Taiwan University, Taipei City.

Pheretima polyglandular  - Chen & Shih, 1996: 54. Fushan Botanical Garden, Ilan County.

Amynthas omeimontis polyglandularis  - Shih et al., 1999: 436.

Amynthas polyglandularis  - Tsai et al., 2000 a: 286; - Chen et al., 2003: 58. Jiaohsi, Ilan County; - Blakemore et al., 2006: 229.

Amynthas omeimontis polyglandular  - Chen et al., 2002: 77. Fushan Botanical Garden, Ilan County; - Chuang et al., 2002: 68. Taipei, Taoyuan and Ilan counties.

Remarks: A. polyglandularis  is considered as A. o. polyglandularis  , a subspecies of Amynthas omeimontis ( Chen, 1931)  by Sims and Easton (1972). However, the arrangement of its genital papillae differs distinctively from that of A. omeimontis  , sufficient for specific distinction ( Tsai et al. 2000 a). Also, A. polyglandularis  has simple intestinal caeca whereas A. omeimontis  has finger-like caeca ( Blakemore et al. 2006).

65. * Amynthas proasacceus  Tsai, Shen & Tsai, 2001

Amynthas proasacceus Tsai et al., 2001: 282  . Mt. Hohuan between Hualien and Nantou counties and Rueyen Creek Nature Reserve, Nantou County; - Tsai et al., 2004 a: 6.

Amynthas assacceus  - Blakemore et al., 2006: 228.

Remarks: A. proasacceus  of Taiwan is considered to be an uncertain synonym of A. asacceus ( Chen, 1938)  of Hainan Island, China, and the latter is the athecate morph of the former ( Blakemore 2003; Blakemore et al. 2006). Both species share fairly similar characteristics in size, large round male porophores, and degenerated spermathecae and male reproductive organs, but they are specifically distinguishable. A. asacceus  has a single seta between the porophores, long coiled prostatic ducts, and male pores, each in a dark-coloured, glandular patch which is 0.7 mm in diameter, ( Chen 1938). A. proasacceus  has 6-9 setae between the porophores, short C-shaped prostatic ducts, and male pores, each in a smooth, indistinctly coloured male patch which is 0.45-0.65 mm in diameter and surrounded by two or three circular folds, (Tsai et al. 2001). Furthermore, recent molecular evidence based on COI sequence analysis demonstrates that A. proasacceus  and A. asacceus  are two valid species ( Chang et al. 2007 a).

66. Amynthas robustus  ( Perrier, 1872)

Pheretima robusta  - Kobayashi, 1940 c: 390. Hsinchu; - Tsai, 1964: 26. The 8 th dormitory of National Taiwan University, Taipei City.

Pheretima lauta Ude, 1905  - Tsai, 1964: 25. National Taiwan University campus and Taipei City Park, Taipei City and Yingko, Taipei County.

Amynthas robustus  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 286; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chen et al., 2002: 77. Fushan Botanical Garden, Ilan County; - Chuang et al., 2002: 68. Taipei, Taoyuan and Hsinchu counties; - Chen et al., 2003: 58. Jiaohsi, Nanao, Fushan Botanical Garden, and Datung, Ilan County; - Tsai et al., 2004 a: 7; - James et al., 2005: 1025. Chaishan, near National Sun Yat-sen University, Kaohsiung City; - Blakemore et al., 2006: 229.

Amynthas lautus  - Tsai et al., 2000 a: 286; - Tsai & Tsai, 2001: 21. National Taiwan University’s Forest Recreation Park, Hsitou, Nantou County; - Shen & Tsai, 2002 b: 3. Lanyu Island.

Amynthas masatakae (Beddard, 1892)  - Chuang & Chen, 2002: 74. Mt. Dongyan, Taoyuan County.

Remarks: A. lautus  is synonymous to A. robustus  according to the recent molecular analysis ( Chang et al. 2007 a). A. masatakae  is also synonymous to A. robustus  according to Easton (1981), Blakemore (2003) and Blakemore et al. (2006).

67. Amynthas rockefelleri  ( Chen, 1933)

Pheretima rockfelleri  - Kobayashi, 1939 a: 660, 1939b: 778, 1940 a: 120, 1940 b: 274, and 1940 c: 391. Hsinchu.

Pheretima rockefelleri  - Tsai, 1964: 8. National Taiwan University campus, Taipei City.

Amynthas hsinpuensis Kuo, 1995: 187  . Hankenli, Hsinpu, Hsinchu County; - Shih et al., 1999: 437; - Tsai et al., 2000 a: 287.

Pheretima rockerfelleri  - Chen & Shih, 1996: 54. Fushan Botanical Garden, Ilan County.

Amynthas papulosus  - Shih et al., 1999: 436 and 437.

Amynthas rockefelleri  - Chen et al., 2003: 58. Nanao, Fushan Botanical Garden and Datung, Ilan County; - Shen et al., 2003 a: 42; - Tsai et al., 2004 a: 6. Meihsi, Wushe and Guohsing, Nantou County.

68. * Amynthas sexpectatus  Tsai, Shen & Tsai, 1999

Amynthas sexpectatus Tsai et al., 1999: 38  . Guohsing, Nantou County; - Tsai et al., 2000 a: 286; - Tsai et al., 2004 a: 6. Jilongshan (Mt. Jilong) near Chungliao, Nantou County; - Blakemore et al., 2006: 229.

69. * Amynthas shinanmontis  Tsai & Shen, 2007

Amynthas shinanmontis Tsai et al., 2007: 358  . Mt. Shinan, Shihshan Forest Road, Chuyunshan Nature Reserve, Kuaigu, and Rt. 20 near Daguanshan Tunnel, Kaohsiung County, and Yakou and Hsiangyang, Taitung County.

70. * Amynthas swanus  ( Tsai, 1964)

Pheretima swanus Tsai, 1964: 13  . National Taiwan University campus, Taipei City.

Amynthas swanus  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 287; - Chuang et al., 2002: 69. Taoyuan County; - Tsai et al., 2004 a: 6. Jiji and Tsaotun, Nantou County; - Blakemore et al., 2006: 229.

71. Amynthas taipeiensis  ( Tsai, 1964)

Pheretima taipeiensis Tsai, 1964: 12  . Chungho, Taipei County; - Chen & Shih, 1996: 54. Fushan Botanical Garden, Ilan County.

Amynthas taipeiensis  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 287; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chuang et al., 2002: 68. Taipei County; - Blakemore et al., 2006: 229.

72. * Amynthas tantulus  Shen, Tsai & Tsai, 2003

Amynthas tantulus Shen et al., 2003 b: 484  . Rueyen Creek Nature Reserve, Nantou County; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 229.

73. * Amynthas tayalis  Tsai, Shen & Tsai, 1999

Amynthas tayalis Tsai et al., 1999: 36  . Hsiju, Taipei County; - Tsai et al., 2000 a: 286; - Chuang et al., 2002: 69. Ilan County; - Chen et al., 2003: 58. Jiaohsi and Datung, Ilan County; - Blakemore et al., 2006: 229.

74. * Amynthas tessellatus tessellatus  Shen, Tsai & Tsai, 2002

Amynthas tessellatus tessellatus Shen et al., 2002: 2  and 7. Wulin to Meihsi, Nantou County; - Tsai, et al., 2004 a: 5; - Blakemore et al., 2006: 229.

75. * Amynthas tessellatus paucus  Shen, Tsai & Tsai, 2002

Amynthas tessellatus paucus Shen et al., 2002: 2  and 7. Wanda to Penpu, Nantou County; - Tsai et al., 2004 a: 5; - Blakemore et al., 2006: 229.

76. * Amynthas tungpuensis  Tsai, Shen & Tsai, 1999

Amynthas tungpuensis Tsai et al., 1999: 34  . Tungpu, Nantou County; - Tsai et al., 2000 a: 286; - Tsai et al., 2004 a: 6. Mountain slops along Penpu, Nanshan and Dongyen creeks, Nantou County; - Blakemore et al., 2006: 229.

Amynthas monsoonus James et al., 2005: 1012  . Mt. Nanren, Pingtung County; - Blakemore et al., 2006: 228.

77. * Amynthas uvaglandularis  Shen, Tsai & Tsai, 2003

Amynthas uvaglandularis Shen et al., 2003 b: 479  . Rueyen Creek Nature Reserve, National Taiwan University’s High-altitude Horticulture Experimental Station at Meifong, and Mt. Beidongyen, Nantou County; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 229.

78. * Amynthas wangi  Shen, Tsai & Tsai, 2003

Amynthas wangi Shen et al., 2003 b: 489  . Rueyen Creek Nature Reserve, Nantou County; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 229.

79. * Amynthas wulinensis  Tsai, Shen & Tsai, 2001

Amynthas wulinensis Tsai et al., 2001: 285  and 2004 a: 5. Wulin to Nanshan Creek, Nantou County; - Blakemore et al., 2006: 229; - Chang et al., 2007 b: 232.

80. * Metaphire bununa  Tsai, Tsai & Liaw, 2000

Metaphire bununa bununa Tsai et al., 2000 c: 1736  . Mt. Jilong, Jiji, Chungliao, Guohsing, and Shuili, Nantou County; - Tsai et al., 2000 a: 287; - Blakemore et al., 2006: 229; - Chang et al., 2008: 958.

Metaphire bununa  - Tsai et al., 2000 a: 286; - Chang & Chen, 2004: 221; - Tsai et al., 2004 a: 5. Mt. Hohuan to Jijji, Nantiu County; - Tsai et al., 2004 b: 881; - Chang & Chen, 2005 a: 593. Heping, Taichung County.

81. Metaphire californica  (Kinberg, 1867)

Pheretima californica  - Kobayashi, 1939 b: 777, 1940a: 120, 1940 b: 274, and 1940 c: 390. Hsinchu; - Gates, 1959: 5. Taipei; - Tsai, 1964: 23. National Taiwan University campus, Taipei City and Yingko, Taipei County; - Kuo, 1995: 185. Tsaochiao, Miaoli County.

Metaphire californica  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 287; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chuang et al., 2002: 68. Taipei, Taoyuan and Hsinchu counties; - Shen & Tsai, 2002 b: 3. Lanyu Island; - Chen et al., 2003: 58. Sanhsing, Tungshan, Yuanshan, Jiaohsi, Nanao, Suao, Datung, and Gueishan (Turtle) Island, Ilan County; - Tsai et al., 2004 a: 6. Coastal plain to the Central Mountain Range to the elevation of 2100 m in central Taiwan; - Chang & Chen, 2005 a: 593. Taipei City; - James et al., 2005: 1026. Laiyi, Pingtung County; - Blakemore et al., 2006: 229.

Metaphire hesperidum (Beddard, 1892)  - Tsai et al., 2000 a: 286.

82. * Metaphire feijani  Chang & Chen, 2004

Metaphire feijani Chang & Chen, 2004: 219  . Majia and Wutai, Pingtung County; - Blakemore et al., 2006: 230; - Chang et al., 2008: 958.

Amynthas ailiaoensis James, Shih & Chang, 2005: 1020  . Wutai, Pingtung County; - Blakemore et al., 2006: 227.

83. * Metaphire formosae  ( Michaelsen, 1922)

Pheretima formosae Michaelsen, 1922: 39  . Chiahsien (= Dorf Koseypo), Kaohsiung County; - Kobayashi, 1939 a: 659. Hsinchu; - Gates, 1959: 9. Chaochou, Pingtung County.

Amynthas formosae  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 286; - Tsai et al., 2000 c: 1740; - Tsai et al., 2004 a: 6. Peripheral hills of Central Mountain Range, Nantou County; - Chang & Chen, 2005 a: 593. Yangmei, Taoyuan County south to Mayshan, Chiayi County; - James et al., 2005: 1017.

Amynthas chaishanensis James et al., 2005: 1021  . Chaishan, Kaohsiung City; - Blakemore et al., 2006: 228.

Metaphire formosae Chang & Chen, 2005 a: 593  . Yangmei, Taoyuan County south to Mayshan, Chiayi County; - Blakemore et al., 2006: 230; - Chang et al., 2008: 958.

84. * Metaphire glareosa  Tsai, Tsai & Liaw, 2000

Metaphire bununa glareosus Tsai et al., 2000 c: 1738  . Jhihben, Taitung County.

Metaphire bununa glareosa  - Blakemore et al., 2006: 229; - Chang et al., 2008: 958.

Metaphire glareosa  - Chang et al., 2008: 964.

85. Metaphire houlleti  ( Perrier, 1872)

Metaphire houlleti  - Shen et al., 2005 a: 13. Haocha, Pingtung County; - James et al., 2005: 1026. Wutai, Pingtung County; - Blakemore et al., 2006: 230.

Amynthas huangi James et al., 2005: 1014  . Shihwen, Pingtung County; - Blakemore et al., 2006: 228.

86. * Metaphire nanaoensis  Chang & Chen, 2005

Metaphire nanaoensis Chang & Chen, 2005 b: 1473  ; - Chen et al., 2003: 58. Nanao, Ilan County; - Blakemore et al., 2006: 230; - Chang et al., 2008: 958.

Metaphire nanauensis  - Chen et al., 2003: 58.

87. * Metaphire paiwanna paiwanna  Tsai, Tsai & Liaw, 2000

Metaphire paiwanna paiwanna Tsai et al., 2000 c: 1732  . Mudan, Pingtung County; - Blakemore et al., 2006: 230; - Chang et al., 2008: 958.

Metaphire paiwanna  - Tsai et al., 2000 a: 286; - Chang & Chen, 2004: 219; - Tsai et al., 2004 b: 881; - Chang & Chen, 2005 a: 593. Liouguei, Kaohsiung County.

Amynthas kaopingensis James et al., 2005: 1017  . Caopu, Shihzih, and Mt. Dahan, Pingtung County; - Blakemore et al., 2006: 228.

88. * Metaphire paiwanna hengchunensis  ( James, Shih & Chang, 2005)

Amynthas hengchunensis James et al., 2005: 1015  . Mt. Nanren and Hengchun, Pingtung County; - Blakemore et al., 2006: 228.

Metaphire hengchunensis  - Chang et al., 2008: 958.

Metaphire paiwanna hengchunensis  - Chang et al., 2008: 965.

89. * Metaphire paiwanna liliumfordi  Tsai, Tsai & Liaw, 2000

Metaphire paiwanna liliumfordi Tsai et al., 2000 c: 1734  . Mt. Setou, Hualien County; - Blakemore et al., 2006: 230; - Chang et al., 2008: 958.

90. Metaphire posthuma  (Vaillant, 1869)

Pheretima posthuma  - Gates, 1959: 15. Chaochou, Pingtung County; - Tsai, 1964: 4. Wanhua railroad station and sand bank of Danshui River, Taipei.

Metaphire posthuma  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 287; - Chuang et al., 2002: 68. Taipei and Ilan counties; - Tsai et al., 2004 a: 6; - Blakemore et al., 2006: 230.

91. * Metaphire puyuma  Tsai, Shen & Tsai, 1999

Metaphire puyuma Tsai et al., 1999: 42  . Monhuan Lake shore, Taitung City; - Tsai et al., 2000 a: 286; - Blakemore et al., 2006: 230.

92. Metaphire schmardae  (Horst, 1883)

Pheretima schmardae  - Kobayashi, 1940 a: 120 and 1940 b: 274. Hsinchu; - Tsai, 1964: 20. National Taiwan University campus and sand bank of Danshui River, Taipei; - Kuo, 1995: 188. Hsinchu Teacher’s College campus.

Metaphire schmardae schmardae  - Shih et al., 1999: 436; - Chang et al., 2001: 76. National Taiwan University campus, Taipei City; - Chang & Chen, 2005 a: 593. Taipei City; - Blakemore et al., 2006: 230.

Metaphire schmardae  - Tsai et al., 2000 a: 288; - Chang et al., 2001: 77. National Taiwan University campus, Taipei City; - Chuang et al., 2002: 68. Taipei and Hsinchu counties; - Tsai et al., 2004 a: 6. Coastal plain to peripheral hills to the elevation of 250 m in central Taiwan.

93. * Metaphire tahanmonta  Chang & Chen, 2005

Metaphire tahanmonta Chang & Chen, 2005 b: 1475  . Mt. Tahan, Pingtung County, and Paoshan and Ernchituan, Kaohsiung County; - Chang & Chen, 2005 a: 593. Taoyuan, Kaohsiung County; - Blakemore et al., 2006: 230; - Chang et al., 2008: 958.

94. * Metaphire taiwanensis  Tsai, Tsai & Shen, 2004

Metaphire taiwanensis Tsai et al., 2004 b: 878  ; - Tsai et al., 2004 a: 6. Mt. Beidongyen and National Taiwan University’s High-Altitude Horticulture Experimental Station, Meifong, Nantou County; - Chang & Chen, 2004: 221; - Blakemore et al., 2006: 230; - Chang et al., 2008: 958.

95. * Metaphire trutina  Tsai, Chen, Tsai & Shen, 2003

Metaphire trutina Tsai et al., 2003: 84  . Hsiaojiaohsi, Ilan County; - Chen et al., 2003: 58. Jiaohsi, Ilan County; - Blakemore et al., 2006: 230; - Chang et al., 2008: 958.

Metaphire yuanpowa Chang & Chen, 2005 b: 1470  . Wulai and Pinglin, Taipei County; - Chang & Chen, 2005 a: 593. Wulai, Taipei County.

96. * Metaphire yeni  Tsai, Shen & Tsai, 2000

Metaphire yeni Tsai S. C. et al., 2000: 8  . Wutai, Pingtung County; - Chang & Chen, 2004: 219; - Blakemore et al., 2006: 230.

97. * Metaphire yuhsii  ( Tsai, 1964)

Pheretima yuhsi Tsai, 1964: 5  . Neihu and Yuantung Temple, Taipei County.

Pheretima formosae  - Kobayashi, 1939 a: 659. Hsinchu; - Gates, 1959: 9. Yangmingshan, Taipei.

Amynthas yuhsi  - Shih et al., 1999: 436.

Amynthas yushi  - Tsai et al., 2000 c: 1740.

Amynthas formosae  - Tsai et al., 2000 a: 286; - Tsai et al., 2000 c: 1740; - Chuang et al., 2002: 68. Taipei and Taoyuan counties; - Chen et al., 2003: 58. Jiaohsi and Gueishan (Turtle) Island, Ilan County; - Chang & Chen, 2005 a: 593.

Metaphire yuhsii Chang & Chen, 2005 a: 593  . Hsintien, Taipei County south to Jianshi, Hsinchu County; - Blakemore et al., 2006: 230; - Chang et al., 2008: 958.

98. Polypheretima elongata  ( Perrier, 1872)

Pheretima elongata  - Gates, 1959: 9. Tsing-Chao Maa, central Taiwan.

Polypheretima elongata  - Shih et al., 1999: 436; - Tsai et al., 2000 a: 288; - Shen & Tsai, 2002 b: 4. Lanyu Island; - Tsai et al., 2004 a: 6. Peripheral hills from the elevation of 250 m to 500 m in central Taiwan; - James et al., 2005: 1026. National Sun Yat-sen University campus, Kaohsiung City; - Blakemore et al., 2006: 230.

COI

University of Coimbra Botany Department

Kingdom

Animalia

Phylum

Annelida

Class

Clitellata

Order

Opisthopora

Family

Megascolecidae

Loc

Megascolecidae

Tsai, Chu-Fa, Shen, Huei-Ping, Tsai, Su-Chen, Lin, Kang-Jieh, Hsieh, Hwey-Lian & Yo, Shao-Pin 2009
2009
Loc

Amynthas nanrenensis James et al., 2005 : 1008

Blakemore 2006: 228
James 2005: 1008
2006
Loc

Amynthas papilio papilio

Blakemore 2006: 228
2006
Loc

Amynthas assacceus

Blakemore 2006: 228
2006
Loc

Amynthas monsoonus James et al., 2005 : 1012

Blakemore 2006: 228
James 2005: 1012
2006
Loc

Amynthas wulinensis Tsai et al., 2001: 285

Chang 2007: 232
Blakemore 2006: 229
2006
Loc

Metaphire feijani

Blakemore 2006: 230
Chang 2004: 219
2006
Loc

Amynthas ailiaoensis

Blakemore 2006: 227
James 2005: 1020
2006
Loc

Amynthas chaishanensis James et al., 2005 : 1021

Blakemore 2006: 228
James 2005: 1021
2006
Loc

Metaphire formosae

Blakemore 2006: 230
Chang 2005: 593
2006
Loc

Metaphire bununa glareosa

Blakemore 2006: 229
2006
Loc

Amynthas huangi James et al., 2005 : 1014

Blakemore 2006: 228
James 2005: 1014
2006
Loc

Metaphire paiwanna paiwanna Tsai et al., 2000 c : 1732

Blakemore 2006: 230
Tsai 2000: 1732
2006
Loc

Amynthas kaopingensis James et al., 2005 : 1017

Blakemore 2006: 228
James 2005: 1017
2006
Loc

Amynthas hengchunensis James et al., 2005 : 1015

Blakemore 2006: 228
James 2005: 1015
2006
Loc

Metaphire paiwanna liliumfordi Tsai et al., 2000 c : 1734

Blakemore 2006: 230
Tsai 2000: 1734
2006
Loc

Metaphire yuhsii

Blakemore 2006: 230
Chang 2005: 593
2006
Loc

Amynthas carnosus

Blakemore 2006: 228
Shen 2005: 97
2005
Loc

Metaphire houlleti

Blakemore 2006: 230
Shen 2005: 13
James 2005: 1026
2005
Loc

Metaphire tahanmonta

Blakemore 2006: 230
Chang 2005: 1475
Chang 2005: 593
2005
Loc

Metaphire yuanpowa

Chang 2005: 1470
Chang 2005: 593
2005
Loc

Amynthas catenus Tsai et al., 2001: 279

Blakemore 2006: 228
Tsai 2004: 6
2004
Loc

Amynthas cortices

Chen 2004: 118
2004
Loc

Amynthas corticis

Blakemore 2006: 228
James 2005: 1023
Tsai 2004: 6
2004
Loc

Amynthas exiguus aquilonius Tsai et al., 2001: 277

Blakemore 2006: 228
Tsai 2004: 6
2004
Loc

Amynthas fenestrus Shen et al., 2003 b : 487

Blakemore 2006: 228
Tsai 2004: 6
Shen 2003: 487
2004
Loc

Amynthas hohuanmontis Tsai et al., 2002: 758

Blakemore 2006: 228
Tsai 2004: 6
2004
Loc

Amynthas nanshanensis Shen et al., 2003 b : 482

Blakemore 2006: 228
Tsai 2004: 6
Shen 2003: 482
2004
Loc

Amynthas penpuensis Shen et al., 2003 b : 481

Blakemore 2006: 228
Tsai 2004: 6
Shen 2003: 481
2004
Loc

Amynthas proasacceus Tsai et al., 2001: 282

Tsai 2004: 6
2004
Loc

Amynthas tantulus Shen et al., 2003 b : 484

Blakemore 2006: 229
Tsai 2004: 6
Shen 2003: 484
2004
Loc

Amynthas tessellatus tessellatus Shen et al., 2002: 2

Blakemore 2006: 229
Tsai 2004: 5
2004
Loc

Amynthas tessellatus paucus Shen et al., 2002: 2

Blakemore 2006: 229
Tsai 2004: 5
2004
Loc

Amynthas uvaglandularis Shen et al., 2003 b : 479

Blakemore 2006: 229
Tsai 2004: 6
Shen 2003: 479
2004
Loc

Amynthas wangi Shen et al., 2003 b : 489

Blakemore 2006: 229
Tsai 2004: 6
Shen 2003: 489
2004
Loc

Metaphire taiwanensis Tsai et al., 2004 b : 878

Blakemore 2006: 230
Tsai 2004: 878
Tsai 2004: 6
Chang 2004: 221
2004
Loc

Metaphire yeni Tsai S. C. et al., 2000: 8

Blakemore 2006: 230
Chang 2004: 219
2004
Loc

Amynthas incongrua

Chen 2003: 58
2003
Loc

Amynthas rockefelleri

Tsai 2004: 6
Chen 2003: 58
Shen 2003: 42
2003
Loc

Metaphire nanaoensis

Blakemore 2006: 230
Chang 2005: 1473
Chen 2003: 58
2003
Loc

Metaphire nanauensis

Chen 2003: 58
2003
Loc

Pithemera lanyuensis

Blakemore 2006: 230
Chang 2004: 219
Shen 2002: 2
2002
Loc

Amynthas aspergillus

Tsai 2004: 5
Chen 2003: 58
Shen 2002: 4
Chang 2001: 76
Tsai 2000: 286
2000
Loc

Amynthas binoculatus Tsai et al., 1999 : 41

Blakemore 2006: 228
Tsai 2004: 6
Tsai 2000: 286
Tsai 1999: 41
2000
Loc

Amynthas polyglandularis

Blakemore 2006: 229
Chen 2003: 58
Tsai 2000: 286
2000
Loc

Amynthas lautus

Shen 2002: 3
Tsai 2000: 286
2000
Loc

Amynthas sexpectatus Tsai et al., 1999 : 38

Blakemore 2006: 229
Tsai 2004: 6
Tsai 2000: 286
Tsai 1999: 38
2000
Loc

Amynthas tayalis Tsai et al., 1999 : 36

Blakemore 2006: 229
Chen 2003: 58
Tsai 2000: 286
Tsai 1999: 36
2000
Loc

Amynthas tungpuensis Tsai et al., 1999 : 34

Blakemore 2006: 229
Tsai 2004: 6
Tsai 2000: 286
Tsai 1999: 34
2000
Loc

Metaphire bununa bununa Tsai et al., 2000 c : 1736

Blakemore 2006: 229
Tsai 2000: 1736
Tsai 2000: 287
2000
Loc

Metaphire bununa

Chang 2005: 593
Chang 2004: 221
Tsai 2004: 5
Tsai 2004: 881
Tsai 2000: 286
2000
Loc

Metaphire hesperidum

Tsai 2000: 286
2000
Loc

Metaphire paiwanna

Chang 2005: 593
Chang 2004: 219
Tsai 2004: 881
Tsai 2000: 286
2000
Loc

Metaphire puyuma Tsai et al., 1999 : 42

Blakemore 2006: 230
Tsai 2000: 286
Tsai 1999: 42
2000
Loc

Metaphire schmardae

Tsai 2004: 6
Chang 2001: 77
Tsai 2000: 288
2000
Loc

Amynthas yushi

Tsai 2000: 1740
2000
Loc

Amynthas formosae

Chang 2005: 593
Chen 2003: 58
Tsai 2000: 286
Tsai 2000: 1740
2000
Loc

Pithemera bicincta

Blakemore 2006: 230
Chen 2003: 58
Shen 2002: 3
Tsai 2000: 288
Shih 1999: 436
1999
Loc

Amynthas aspergillum

Blakemore 2006: 227
Shih 1999: 436
1999
Loc

Amynthas candidus

Blakemore 2006: 228
Tsai 2000: 286
Shih 1999: 436
1999
Loc

Amynthas corticus

Chen 2003: 58
Shen 2002: 3
Chang 2001: 76
Tsai 2000: 286
Shih 1999: 436
1999
Loc

Amynthas gracilis

Blakemore 2006: 228
James 2005: 1024
Chen 2004: 118
Tsai 2004: 6
Chen 2003: 58
Shen 2002: 3
Chang 2001: 76
Tsai 2000: 286
Shih 1999: 436
1999
Loc

Amynthas hupeiensis

Blakemore 2006: 228
Tsai 2004: 6
Chang 2001: 76
Tsai 2000: 287
Shih 1999: 436
1999
Loc

Amynthas incongruus

Blakemore 2006: 228
James 2005: 1025
Tsai 2004: 6
Tsai 2000: 287
Shih 1999: 436
1999
Loc

Amynthas minimus

Blakemore 2006: 228
Tsai 2000: 287
Shih 1999: 436
1999
Loc

Amynthas morrisi

Blakemore 2006: 228
Tsai 2004: 6
Tsai 2000: 287
Shih 1999: 436
1999
Loc

Amynthas papulosus

Blakemore 2006: 228
Tsai 2004: 6
Chen 2003: 58
Chang 2001: 76
Tsai 2000: 287
Shih 1999: 436
1999
Loc

Amynthas omeimontis polyglandularis

Shih 1999: 436
1999
Loc

Amynthas robustus

Blakemore 2006: 229
James 2005: 1025
Tsai 2004: 7
Chen 2003: 58
Chang 2001: 76
Tsai 2000: 286
Shih 1999: 436
1999
Loc

Amynthas hsinpuensis

Tsai 2000: 287
Shih 1999: 437
Kuo 1995: 187
1999
Loc

Amynthas papulosus

Shih 1999: 436
1999
Loc

Amynthas swanus

Blakemore 2006: 229
Tsai 2004: 6
Tsai 2000: 287
Shih 1999: 436
1999
Loc

Amynthas taipeiensis

Blakemore 2006: 229
Chang 2001: 76
Tsai 2000: 287
Shih 1999: 436
1999
Loc

Metaphire californica

Blakemore 2006: 229
Chang 2005: 593
James 2005: 1026
Tsai 2004: 6
Chen 2003: 58
Shen 2002: 3
Chang 2001: 76
Tsai 2000: 287
Shih 1999: 436
1999
Loc

Amynthas formosae

Chang 2005: 593
James 2005: 1017
Tsai 2004: 6
Tsai 2000: 286
Tsai 2000: 1740
Shih 1999: 436
1999
Loc

Metaphire posthuma

Blakemore 2006: 230
Tsai 2004: 6
Tsai 2000: 287
Shih 1999: 436
1999
Loc

Metaphire schmardae schmardae

Blakemore 2006: 230
Chang 2005: 593
Chang 2001: 76
Shih 1999: 436
1999
Loc

Amynthas yuhsi

Shih 1999: 436
1999
Loc

Polypheretima elongata

Blakemore 2006: 230
James 2005: 1026
Tsai 2004: 6
Shen 2002: 4
Tsai 2000: 288
Shih 1999: 436
1999
Loc

Pheretima polyglandular

Chen 1996: 54
1996
Loc

Pheretima rockerfelleri

Chen 1996: 54
1996
Loc

Pheretima taipeiensis

Chen 1996: 54
Tsai 1964: 12
1996
Loc

Pheretima lauta

Tsai 1964: 25
1964
Loc

Pheretima rockefelleri

Tsai 1964: 8
1964
Loc

Pheretima bicincta

Gates 1959: 4
1959
Loc

Pheretima aspergillum

Tsai 1964: 29
Gates 1959: 1
1959
Loc

Pheretima candida

Gates 1959: 18
1959
Loc

Pheretima incongrua

Tsai 1964: 19
Gates 1959: 13
1959
Loc

Pheretima papulosa

Gates 1959: 18
1959
Loc

Pheretima posthuma

Tsai 1964: 4
Gates 1959: 15
1959
Loc

Pheretima elongata

Gates 1959: 9
1959
Loc

Pheretima zoysiae

Kobayashi 1941: 378
Kobayashi 1940: 274
1940
Loc

Pheretima robusta

Tsai 1964: 26
Kobayashi 1940: 390
1940
Loc

Pheretima schmardae

Kuo 1995: 188
Tsai 1964: 20
Kobayashi 1940: 120
1940
Loc

Pheretima takatorii

Kobayashi 1939: 659
1939
Loc

Pheretima diffringens

Chen 1996: 54
Tsai 1964: 2
Gates 1959: 6
Kobayashi 1939: 660
1939
Loc

Pheretima hawayana

Chen 1996: 54
Kuo 1995: 184
Tsai 1964: 9
Kobayashi 1939: 778
1939
Loc

Pheretima hupeiensis

Kobayashi 1939: 660
1939
Loc

Pheretima morrisi

Tsai 1964: 17
Kobayashi 1939: 778
1939
Loc

Pheretima papulosa sauteri

Kobayashi 1939: 660
Michaelsen 1922: 26
1939
Loc

Pheretima rockfelleri

Kobayashi 1939: 660
1939
Loc

Pheretima californica

Kuo 1995: 185
Tsai 1964: 23
Gates 1959: 5
Kobayashi 1939: 777
1939
Loc

Pheretima formosae

Gates 1959: 9
Kobayashi 1939: 659
Michaelsen 1922: 39
1939
Loc

Pheretima formosae

Gates 1959: 9
Kobayashi 1939: 659
1939
Loc

Perionyx excavatus

Blakemore 2006: 230
Tsai 2004: 6
Chang 2001: 76
Tsai 2000: 286
Shih 1999: 436
Kuo 1995: 189
Gates 1959: 19
Kobayashi 1938: 201
1938
Loc

Pheretima

Takahashi 1931: 10
1931
Loc

Pheretima (Pheretima) aspergillum

Michaelsen 1929: 158
1929
Loc

Perichaeta takatorii

Goto 1898: 76
Loc

Amynthas bilineatus Tsai et al., 2007 : 366

Tsai 2007: 366
Loc

Perichaeta candida

Goto 1898: 77
Loc

Amynthas chilanensis Tsai et al., 2007 : 362

Tsai 2007: 362
Loc

Amynthas cruxus Tsai et al., 2007 : 368

Tsai 2007: 368
Loc

Amynthas lini Chang et al., 2007 b : 234

Chang 2007: 234
Loc

Amynthas meishanensis Chang et al., 2007 b : 236

Chang 2007: 236
Loc

Pheretima polyglandularis

Tsai 1964: 30
Loc

Amynthas shinanmontis Tsai et al., 2007 : 358

Tsai 2007: 358
Loc

Pheretima swanus

Tsai 1964: 13
Loc

Metaphire bununa glareosus Tsai et al., 2000 c : 1738

Tsai 2000: 1738
Loc

Pheretima yuhsi

Tsai 1964: 5