Cyclocephala carbonaria Arrow, 1911

Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 165

§ Cyclocephala carbonaria Arrow, 1911

Cyclocephala carbonaria Arrow 1911b: 173 ( Nicaragua, Panama)

Distribution. Venezuela (EO): Joly 1995: 171 (EO); Moore et al. 2018b: 156 (EO). Other: Arrow 1911b: 173 (Nicaragua, Panama), 1937b: 9 (Ecuador, Nicaragua, Panama); Blackwelder 1944: 251 (Ecuador, Nicaragua, Panama); Endrödi 1966a: 164 (Bolivia, Colombia, Costa Rica, Ecuador, Mexico, Nicaragua, Panama), 1985a: 40, 117 (Bolivia, Costa Rica, Ecuador, Mexico, Nicaragua, Panama); Joly 1995: 171 (Colombia, Costa Rica, Ecuador, Nicaragua, Panama); Ratcliffe 2003: 99 (Costa Rica, Honduras, Mexico, Panama); Moore et al. 2018b: 156 (Belize, Bolivia, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama).

Note. Dechambre (1997a) splits the species Cyclocephala carbonaria into a number of species, with C. carbonaria itself being restricted to Central America. According to this, South American specimens previously identified as C. carbonaria , such as those mentioned for Venezuela in Joly (1995: 171) and Moore et al. (2018b: 156) (the latter who simply copied from older publications), must thus belong to other species. Ratcliffe (2003: 99) suggests that more research needs to be done to establish if the South American specimens actually represent separate species or if they are merely intraspecific variations within C. carbonaria . The taxa which are described by Dechambre (1997a) and which are known to occur in the research area are C. boucheri and C. ergastuli . These are considered valid species by us and are included in this checklist.