Sagola bituberata Broun, 1914

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 106-107

Sagola bituberata Broun, 1914

( Figs. 58b , 59b , 60 )

Sagola bituberata Broun, 1914b: 160 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.

Type Material. One of three syntypes. NEW ZEALAND: Buller : 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “ 3525. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Greymouth. J.H. Lewis ” [white label, handwritten]; “ Sagola ♂. bituberata” [white label, handwritten] .

Additional Material (n = 52: 24 males; 28 females). NEW ZEALAND: Buller : 3♂♂, Lake nr Mt. Priestly, Paparoa Ra, 1060m, 10 XII 1969, J. McBurney, litter; 1♂, Fletchers Ck, 6km sw of Rotokohu , 25 I 1972, J.M. McBurney, litter ; 1♀, Relax Shelter, Mt Robert Ridge , 1 X 2005, J. Nunn, moist tussock litter at 1410m ; 1♂, Arthur Pass, Kellys Ck , 8 XI 2005, R. Leschen, S. Nomura, litter, 42′47S, 171′24E; Nelson : 2♂♂ 3♀♀, Canaan , 18 IV 1966, J.I. Townsend, litter ; 1♂, Mt. Domett , 1250m, 30 XI 1971, G. Kuschel, litter ; 2♀♀, Canaan, Harwoods tr, 4 II 1965, L.P. Marchant, litter; Otago Lakes : 1♂ 1♀, Haast River, 100m, Sunny Flat , 25 I 1978, G. Kuschel, litter ; 2♀♀, Mt. Aspiring NP, 5.5km, 11–17 I 1985, Nothofagus menziesii forest, A. Newton, M. Thayer 741, window trap; Westland : 4♂♂ 8♀♀, Kellys Creek, Otira , 22 III 2009, J. Nunn, mossy forest floor litter ; 1♂ 2♀♀, Kellys Creek, Otira , 25 II 1989, J. Nunn ; 1♀, Haast Pass n of summit, 550m, 24 I 1978, G. Kuschel, sifted litter and rotten wood ; 1♂ 2♀♀, Okuku Ck, 11.3km sse Kumara , 60m, 18–22 III 1980, podocarpbroadlf, A. Newton, M. Thayer, litter ; 3♂♂, Franz Joseph. 24 II 1989, J. Nunn ; 2♀♀, Lake Wombat tr, Franz Joseph , 7 II 2009, J. Nunn, washed soil sample, tree fern/broadleaf forest ; 2♂♂, Jackson Bay, Smoothwater tr, 14 II 1977, T. K. Crosby, litter ; 1♂, Kellys Cr, Otira Vly , 9 IV 2010, J. Nunn, litter in kamahi forest ; 1♂, 2.7km s Franz Josef Glacier, #063, Trailhead of Alex Knob , 43′28S 1173′10E, 150m, coastal forest litter, 15 I 1998, C. Carlton, R. Leschen ; 1♂, Mt. Aspiring NP, 5.5km nne Makarora , 330m, 11–17 I 1985, Nothofagus menziesii forest, A. Newton, M. Thayer 741, FIT & window trap ; 1♂, Mt. Aspiring NP, Douglas Creek at Hwy. 6, 65m, 11–18 I 1985, Nothofagus menziesii -hdwd-podocarp forest, A. Newton, M. Thayer 738, FIT & window trap ; 1♀, Open Bay I, Taumaka I, 27 IX 1976, D.S. Horning, litter ; 1♀, Mt Tuhua , 1067m, e side of L. Kaniere, 20 XI 1984, C.F. Butcher, litter ; 1♀, Lake Ianthe , 7 II 1984, J.C. Watt, sifted litter and wood mould ; 1♀, Doughboy Creek, 6km sw Mahitahi , 5 II 1984, J.C. Watt, sifted wood mould .

Diagnosis. This species can be distinguished from the other species of the misella speciesgroup by the following combination of characters: body larger, length 2.5–3.1 mm; male antennomeres 4–10 enlarged; male abdominal ventrite V with weak process medially; shape of antennomeres and genitalia unique to species.

Redescription. Length 2.5–3.1 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 58b ). Head: Round, as long as wide, widest across eyes ( Fig. 58b ). Male antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate, 4–10 enlarged, 5–11 with tubercles. Female antennomeres 2–8 longer than wide, 9–10 subquadrate, 5–11 with tubercles. Frontal rostrum rectangular, lobes continuous. Frontal sulcus reaching midpoint of eye. Posterior frontal fovea teardrop-shaped. Eye prominent, approximately one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Male elytra rectangular ( Fig. 58b ), female subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia, patches absent in female. Male abdominal ventrite V with weak process medially. Aedeagus: Apical lobe of genitalia arrowhead-shaped ( Fig. 59b ). Phallobase of median lobe symmetrical and rounded ( Fig. 59b ). Parameres asymmetrical, right paramere longer than left with setae apically ( Fig. 59b ).

Type Locality. Greymouth , Buller, New Zealand .

Distribution. Buller, Nelson, Otago Lakes, Westland ( Fig. 60 : triangles).

Habitat. Specimens of this species were collected mostly by sifting leaf litter, and a few male specimens were from flight intercept or window traps.

Figures

Fig. 58. Habitus of Sagola species in the misella species-group, dorsal views. a) S. misella, b) S. bituberata, c) S. prisca, d) S. murchisonensis, e) S. fiordlandensis, f) S. hunterensis, g) S. humpridgensis, h) S. clunieae, i) S. turretensis, j) S. otagoensis, k) S. southlandensis, l) S. huvali, m) S. tennysonensis. Scale bars = 1 mm.

Fig. 59. Sagola species in the misella species-group. Aedeagi, dorsal views: a) S. misella, b) S. bituberata, c) S. prisca, d) S. murchisonensis, e) S. fiordlandensis, f) S. hunterensis, g) S. humpridgensis, h) S. clunieae, i) S. turretensis, j) S. otagoensis, k) S. otagoensis, apical lobe, lateral view, l) S. southlandensis, m) S. huvali, n) S. tennysonensis. S. misella: o) Dorsal surface of head, p) Prosternum, ventral view, q) Meso- and metaventrites, ventral view, r) Abdomen, ventral view. Scale bars = 0.1 mm.

Fig. 60. Known collection localities of New Zealand Sagola species. S. misella: black circles; S. bituberata: triangles.