Sagola notabilis Broun, 1880, 2013

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 98-100

Sagola notabilis Broun, 1880

( Figs. 56a, 56d , 57 )

Sagola notabilis Broun, 1880: 137 . Schaufuss 1888: 85; Raffray 1893: 32, 1904: 496, 1911: 5, 1924: 231; Hudson 1923: 365, 1934: 183; Kuschel 1990: 48; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.

Sagola macronyx Broun, 1893b: 1418 . Raffray 1904: 497, 1911: 5, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243. New synonymy.

Sagola electa Broun, 1914a: 91 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.

Sagola fulvipennis Broun, 1915: 289 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.

Type Material. Lectotype. NEW ZEALAND: Coromandel : 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “251” [white label, printed]; “Tairua” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola notabilis ” [white label, handwritten]; “ LECTOTYPE Sagola notabilis Broun Desig. Park and Carlton 2013 ” [red label, printed] . Paralectotype (1 male). NEW ZEALAND: Coromandel : 1♂ ( BMNH) , Tairua, New Zealand, Sharp Coll. 1905-313 ( BMNH). The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that two specimens were examined (Broun 1880: 137). This designation will fix the identity of the species and facilitate its recognition by future workers. Holotype of Sagola electa : NEW ZEALAND: Taupo : 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “3400. ♂ ” [white label, handwritten]; “Erua. Jany.1911.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola electa . ♂.” [white label, handwritten] . Holotype of Sagola fulvipennis : NEW ZEALAND: Auckland: 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “3705. ♂ ” [white label, handwritten]; “Tairua. Novr.1875.” [white label, handwritten]; “ Sagola ♂. fulvipennis” [white label, handwritten]. The original description states that the specimen was collected in December 1874, but the single specimen examined was collected in November 1875. Perhaps the date provided in the original description was incorrect . Holotype of Sagola macronyx : NEW ZEALAND: Auckland: 1♂ ( BMNH), glued on rectangular card, “ Type ” [red label, printed]; “ ♂ ” [white label, printed]; “977” [white label, printed]; “Hunua” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola macronyx ” [white label, handwritten] .

Additional Material (n = 72: 27 males; 45 females). NE W Z EAL AND: Au ck la nd: 23♂♂ 34♀♀, Lynfield, G. Kuschel, 23 XI 1974 – 29 III 1981, malaise trap or litter; 2♂♂, Waitakere Range, 260m, Nohoanga Scenic Res, 8 XII 1984 – 25 I 1985, hdwd-podocarp forest, A. Newton, M. Thayer 679, FIT &window trap; 1♀, Hunua Ranges, Vining Res, Mangatangi tr, 240m, 37″07,747′S, 175″13.024′E, 17 XI–28 XII 2005, Nothofagus - Agathis -Phyllocladus trichomanoides, etc, FMHD #2005-005, FIT, A. Newton, M. Thayer et al, ANMT site 1141; 2♀♀, Orere Saddle, 2 I 1984, J.C. Watt, bark and subcortical debris 84/1; 1♀, Manurewa, Murphy’ s Bush, 2 III 1981, G. Kuschel, rotten wood 81/59; 1♀, Papatoetoe, Murphy’ s Bush, 17 VIII 1978, D.W. Helmore, woodmould 78/204; Bay of Plenty: 1♂, Mt Te Aroha Summit, 1000m, 3 XI 1977, B.M. May, dead trunk; 1♂, Te Aroha, 19 III 1975, G. Kuschel, sifted rotten wood bottom stream; 1♀, Okere Falls Scenic Res, 21 X 1979, J.S. Dugdale, liverworts 79/99; Gisborne: 2♀♀, Gray’ s Bush, 30 VII 1951, under bark; Northland: 2♀♀, Omahuta Kauri Sanctuary, 4 II 1975, G. Kuschel, rotten wood; Rangitikei: 1♀, Ruahine Range, Armstrong Saddle, 10 II 1980, C.F. Butcher, litter 80/11.

Diagnosis. This species can be distinguished from the other species of the notabilis speciesgroup by the following combination of characters: body length 2.4–2.7 mm; male antennomere 8 enlarged; male fore femur with weak semicircular depression; shape of ventral surface of male head and genitalia unique to species; only known from North Island.

Redescription. Length 2.4–2.7 mm. Body brown, antennae, legs, maxillary palpi,and elytra paler ( Fig. 56a ). Head: Round, as long as wide, widest across eyes ( Fig. 56a ). Antennomere 2 longer than wide, 3 subquadrate, 4–7 longer than wide, 5–10 subquadrate, 8 enlarged in male. Eye prominent, approximately two-thirds length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular ( Fig. 56a ). Meso- and metathorax trapezoidal, longer than wide. Male fore femur with shallow, semicircular depression. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe of genitalia triangular and shorter than parameres ( Fig. 56d ). Phallobase of median lobe symmetrical and rectangular ( Fig. 56d ). Parameres symmetrical, setose from midpoints to apices ( Fig. 56d ).

Type Locality. Tairua , Coromandel, New Zealand .

Distribution. Auckland, Bay of Plenty, Coromandel, Gisborne, Northland, Rangitikei ( Fig. 57 : black circles).

Habitat. Specimens of this species were collected using Malaise, flight intercept, and window traps or by sifting dead logs and wood litter in hardwood, podocarp, or Nothofagus forests.

Comments. Male specimens of S. notabilis can be distinguished from those of other species by the enlarged male antennomere 8 and modification of the ventral surface of the male head. The type specimens of S. electa , S. fulvipennis , and S. macronyx share these diagnostic characters, and these species have been collected at or near the type locality. For these reasons, we have placed S. electa , S. fulvipennis , and S. macronyx in synonymy with S. notabilis .

Figures

Fig. 56. Sagola species in the notabilis species-group. Habitus, dorsal views: a) S. notabilis, b) S. monstrosa, c) S. tararuaensis. Aedeagi, dorsal views: d) S. notabilis, e) S. monstrosa, f) S. tararuaensis. S. monstrosa, male: g) Antenna, h) Dorsal surface of head, i) Ventral surface of head, j) Hind leg, lateral view, k) Hind claw, lateral view, l) Abdomen, ventral view. Scale bars a–c = 1 mm, d–l = 0.1 mm.

Fig. 57. Known collection localities of New Zealand Sagola species. S. notabilis: black circles; S. monstrosa: triangles; S. tararuaensis: black squares.