Sagola insignis, Broun, 1893 b:, 1049

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 123-124

28. insignis species-group

(24 species)

Diagnosis. Members of the insignis speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.2–3.2 mm; antennomere 1 approximately 1.5 times longer than wide; male head triangular, widest across temples ( Fig. 70y–z ) and female head bluntly triangular, widest across eyes; anterior and posterior frontal fovea present ( Fig. 70y–z ); temples of male head depressed and projecting ventrally ( Fig. 69 ); male neck ventrally with patch of long and dense setae at anteriorly ( Fig. 69 ); female hind wings reduced to small pads; abdominal tergites IV–VI with discal carinae; genitalia broad and flattened ( Fig. 70a–x ).

KEY TO SPECIES OF THE INSIGNIS SPECIES- GROUP

The key is based on male specimens because most female specimens are indistinguishable based on external morphology.

1. Anterior margin of temporal lobe depressed and L-shaped ( Fig. 69b, s ) ..... ............................................................... 2

1′. Anterior margin of temporal lobe straight or weakly curved...................................3

2(1). Apical lobe of genitalia triangular ( Fig. 70b ); known from South Island ( Fig. 71 : triangles) ................................ ................................. S. hectorii Broun

2′. Apical lobe of genitalia round ( Fig. 70s ); known from North Island ( Fig. 74 : black circles) ...... S. hanae new species

3(1′). Temporal lobe of head weakly projecting, as long as eye or barely exceeding eye ( Fig. 69c, f–g, k )..........................4

3′. Temporal lobe of head distinctly projecting, obviously exceeding eye ( Fig. 69a– b, d–e, h–j, l–x )..................................... 7

4(3). Median lobe of genitalia with a pair of processes on lateral margins...............5

4′. Median lobe of genitalia without processes on lateral margins.....................6

5(4). Apical lobe of genitalia longer than wide ( Fig. 70c )..... S. bipunctata Broun

5′. Apical lobe of genitalia subquadrate ( Fig. 70f )............... S. robustula Broun

6(4′). Elytra rectangular; median lobe of genitalia widest at midpoint ( Fig. 70g )....... ................................ S. pertinax Broun

6′. Elytra subquadrate; median lobe of genitalia widest at one-third length ( Fig. 70k ) ..... S. sheldoni new species

7(3′). Elytra rectangular................................8

7′. Elytra subquadrate.............................18

8(7). Median lobe of genitalia with pair of acute processes laterally......................9

8′. Median lobe of genitalia without process ............................................................. 14

9(8). Apical lobe of genitalia with round depression at apex.............................10

9′. Apical lobe of genitalia with round or straight apex ...................................... 11

10(9). Apical lobe of genitalia semicircular ( Fig. 70a ) .................. S. insignis Broun

10′. Apical lobe of genitalia subquadrate ( Fig. 70d )............. S. angulifera Broun

11(9′). Pair of acute processes on lateral margin of genitalia reaching apex of apical lobe ( Fig. 70m ) ......................... ................. S. cuvierensis new species

11′. Pair of acute processes on lateral margin of genitalia shorter than apex of apical lobe....................................................12

12(11′). Pair of acute processes on lateral margin of genitalia larger, reaching one-third to one- fourth length of apical lobe ....... 13

12′. Pair of acute processes on lateral margin of genitalia minute, not exceeding onefifth length of apical lobe ( Fig. 70q ) .... ............... S. boudreauxae new species

13(12). Median lobe at least three times broader than parameres ( Fig. 70e ); known from North Island ( Fig. 71 : white circles) .... ................................. S. eminens Broun

13′. Median lobe approximately twice broader than parameres ( Fig. 70i ); known from South Island ( Fig. 72 : black squares) .... .................................... S. laticeps Broun

14(8′). Parameres with more than 10 setae apically ( Fig. 70j )........ S. castanea Broun

14′. Parameres with less than 5 setae apically...................................................... 15

15(14′). Apical lobe of genitalia with semicircular depression apically ( Fig. 70r ) .......... ......................... S. taegyui new species

15′. Apical lobe of genitalia without depression apically.......................................16

16(15′). Apical lobe of genitalia rectangular ( Fig. 70l )................................................ ............... S. mercuryensis new species

16′. Apical lobe of genitalia triangular....17

17(16′). Median lobe of genitalia widest at midpoint ( Fig. 70h ) .... S. monticola Broun

17′. Median lobe of genitalia widest at base ( Fig. 70u )............................................... ................. S. egmontensis new species

18(7′). Apical lobe of genitalia with acute apex................................................................19

18′. Apical lobe of genitalia with blunt apex...........................................................20

19(18). Median lobe of genitalia with pair of acute processes laterally ( Fig. 70n ) ...... ....................... S. buckleyi new species

19′. Median lobe of genitalia without processes laterally ( Fig. 70p ) ..................... ............................ S. otteai new species

20(18′). Parameres triangular ( Fig. 70t ) ............. ...................... S. gyudongi new species

20′. Parameres slender..............................21

21(20′). Apical lobe of genitalia triangular....22

21′. Apical lobe of genitalia rectangular .... 23

22(21). Median lobe of genitalia widest at onethird length ( Fig. 70x ); known from South Island ( Fig. 74 : white squares) ... ................. S. jeongnamae new species

22′. Median lobe of genitalia widest at midpoint ( Fig. 70o ); known from North Island ( Fig. 73 : black squares).............. .................... S. brumfieldi new species

23(21′) Median lobe of genitalia widest at onethird length ( Fig. 70w ); known from Stewart Island ( Fig. 74 : white circles) .............. S. stewartensis new species

23′. Median lobe of genitalia widest at base ( Fig. 70v ); known from North Island ( Fig. 74 : stars) ..... S. boonei new species

Figures

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 71. Known collection localities of New Zealand Sagola species. S. insignis: black circles; S. hectorii: triangles; S. bipunctata: black squares; S. angulifera: stars; S. eminens: white circles; S. robustula: white squares.

Fig. 74. Known collection localities of New Zealand Sagola species. S. hanae: black circles; S. gyudongi: triangles; S. egmontensis: black squares; S. boonei: stars; S. stewartensis: white circles; S. jeongnamae: white squares.

Fig. 72. Known collection localities of New Zealand Sagola species. S. pertinax: black circles; S. monticola: triangles; S. laticeps: black squares; S. castanea: stars; S. sheldoni: white circles; S. mercuryensis: white squares.

Fig. 73. Known collection localities of New Zealand Sagola species. S. cuvierensis: black circle; S. buckleyi: triangle; S. brumfieldi: black squares; S. otteai: stars; S. boudreauxae: white circle; S. taegyui: white squares.