Sagola strialis Broun, 1921

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 74-78

Sagola strialis Broun, 1921

( Figs. 43–44 )

Sagola strialis Broun, 1921a: 489 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.

Sagola fasciculata Broun, 1921a: 497 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.

Type Material. Holotype. NEW ZEALAND: Mid Canterbury: 1♂ ( BMNH), glued on rectangular card, “3996. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Moa Hill. 20.11.1913.” [white label, handwritten]; “ Sagola . ♂ strialis.” [white label, handwritten] . Syntypes. NEW ZEALAND: Nelson: 1♂ ( BMNH), glued on rectangular card, “4007.♂” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Glenhope. 20.12.1914 ” [white label, handwritten]; “ Sagola fasciculata ♂.”[white label, handwritten]. 1♂ ( BMNH), glued on rectangular card, “4007. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Glenhope. 20.2.1915 ” [white label, handwritten]; “ Sagola ♂ fasciculata.”[white label, handwritten]. 1♀ ( BMNH), glued on rectangular card, “4007. ♀ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Glenhope. 20.10.1914 ” [white label, ♂♂ handwritten]; “ Sagola ♀ fasciculata.” [white label, handwritten] .

Additional Material (n = 246: 160 males; 86 females). NEW ZEALAND: Buller: 13♂♂ 3♀♀, Nelson Lakes NP, Mt. Robert, Speargrass tr, 875m, 41″49.469′S, 172″48.311′E. 17 XII 2005, Nothofagus forest, FMHD #2005-110, berl leaf & log litter, A. Solodovnikov, D. Clarke, ANMT site 1161; 1♂, Nelson Lakes NP, Mt. Robert, Speargrass tr, 875m, 41″49.469′S, 172″48.311′E. 30 XI–17 XII 2005, Nothofagus forest, FMHD #2005-061, pitfall trap (10), A. Solodovnikov, D. Clarke, ANMT site 1161; 1♂, Nelson Lakes NP, Mt. Robert, Speargrass tr, 875m, 41″49.469′S, 172″48.311′E. 30 XI 2005, Nothofagus forest, FMHD #2005-110, fogging, A. Newton, ANMT site 1161; 7♂♂ 3♀♀, Lewis Pass Nat Res, 11.9km ese Springs Junction, 540m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 715, litter; 3♂♂, 18km n Punakaiki, 80m, 19 XII 1984 – 20 I 1985, hdwd forest with nikau, A. Newton, M. Thayer 718, litter; 3♂♂, Nelson Lks NP, Lake Rotoroa, Braeburn tr, 470m, 16 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 712, window trap; 4♂♂ 2♀♀, Nelson Lks NP, n slope Mt. Robert, Pinchgut tr, 950m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 707, litter; 1♂, Lewis Pass NR, 13.2km s Lewis Pass, 650m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 713; 3♂♂ 4♀♀, Lewis Pass NR, 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, litter; 2♂♂ 3♀♀, Nelson Lks NP, n slope Mt. Robert, 860m, 23–26 III 1980, Nothofagus forest, A. Newton, M. Thayer, litter; 2♂♂, 0.8km n Bullock Ck nr Punakaiki, 50m, 23 III 1980, broadleaf-nikau palm-podocp, A. Newton, M. Thayer, litter; 1♂, Nelson Lks NP, Lake Rotoiti, St. Arnaud tr, 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &window trap; 1♂, Greymouth, Boddytown, 2 II 1984, J.C. Watt, sifted litter 84/23; 1♀, Paparoa Ra, 610m, Buckland Peaks Tk, 25 XI 1984, B.A. Holloway, C.F. Butcher, moss & lichen 84/77; 3♂♂, Flagstaff Bio. Res., 13 VII 1972, J.S. Dugdale, litter; 1♂, Mawhera SF, 15km se of Ngahere, 305m, 10 XI 1971, J.S. Dugdale, litter 71/143; 1♂, Mawhera SF, 19km se of Ngahere, 305m, 10 XI 1971, J. McBurney, litter 71/144; 1♂, Mawhera SF, 16km se of Ngahere, 305m, 11 XI 1971, J.S. Dugdale, litter 71/145; 2♂♂ 1♀, Mawhera SF, Kangaroo Ck, 27 I 1972, J.S. Dugdale, litter 72/86; 2♂♂, Paparoa Ra, Croesus tr, 730m, 21 I 1982, J.W. Early, moss & litter in podocarp –broadleaf forest; 1♂ 1♀, Fletchers Ck, Biological Reserve, XI 1971, McBurney, litter 71/127; 1♂, W. Inangahua, St. 127 Coll Ck, 19 IX 1972, J.S. Dugdale, moss 72/184; 1♂, Tawhai SF, Big R. Rd, 3km S of Reefton, 28 I 1972, J. McBurney, litter 72/76; 1♂, Fletcher Ck, MacMahan Ck, 25 I 1972, J.S. Dugdale, XPB9, litter 72/79; 1♂, Fletchers Ck, 6km sw Rotokohu, 25 I 1972, J.S. Dugdale, PB1 litter 72/103; 1♂, Fletchers Ck, 6km sw Rotokohu, 25 I 1972, J. McBurney, PB15 litter 72/93; 1♂, Capleston, Redmans Ck, 21 IX 1972, J.C. Watt, litter 72/186; 1♂, Reefton, 12 IV 1977, J.A. Wightman, pit trap burnt pine; 1♂, Lewis Pass, 19 X 2007, J. Nunn, washed soil sample; 1♂, Inland Pack tr, 1km n of Runakaiki River, sifted moss, J. Nunn, 29 XII 2010; 1♀, Coal Ck, Runanga, 11 II 2007, J. Nunn, moss; 1♀, W Inangahua SF, 127m, Stoney Ck, 18 IV 1972, J. McBurney, litter PB1; 1♀, Fletcher Ck, 7 III 1972, J.S. Dugdale, litter 72/106; 1♀, Reefton, 5 XI 1958, J.M. Hoy; 1♂, Nelson Lks NP, Mt. Robert car park, 28 IX 1973, R.M. Emberson, beech litter; Fiordland: 1♂ 1♀, W. Olivine Ra, Simonin Pass, 1 II 1975, G.W. Ramsay, under decayed branches lower bush edge; 2♂♂ 1♀, Fiordland NP, Borland rd, Borland Saddle vic, 980m, 45″44.81′S 167″22.88′E, 10 XII 2005, Nothofagus menziesii forest in ravine, FMHD #2005-090, litter, A. Newton, ANMT site 1171; 1♂ 4♀♀, Fiordland NP, Milford Sound rd, Tukoko tr, 70m, 44″40.517′S 167″57.895′E, 9 XII 2005, very mossy Nothofagus forest, emergent podocarp, FMHD #2005-086, litter, A. Newton, M. Thayer, ANMT site 1168; 1♂ 1♀, Fiordland NP, Milford Sound rd, Gertrude Saddle car park area, 800m, 44″46.13′S 168″00.16′E, 9 XII 2005, very mossy Nothofagus menziesii forest, FMHD #2005-087, litter, A. Newton, M. Thayer, ANMT site 1169; 1♂ 6♀♀, Fiordland NP, Milford Sound rd, Smithy Creek Campground area, 400m, 44″57.065′S 168″01.155′E, 9 XII 2005, Nothofagus fusca & N. menziesii open forest, FMHD #2005-089, litter, M. Thayer, A. Newton, ANMT site 1170; 1♂, Borland rd, Limestone Cave tr, 45′46S 167′28E, #140, Nothofagus log/bark litter berlese, 24 I 1998, C. Carlton, R. Leschen; 1♂, Borland rd, Borland Saddle tr, 45′46S 167′23E, #138, Nothofagus leaf litter berlese, 24 I 1998, C. Carlton, R. Leschen; 1♂, Bauza Is, Doubtfull sd, III 1986, pitfall trap, G.M. Bremner; 1♂, Lower Hollyford rd, 44′44S 168′08E, #111, Nothofagus /coastal forest leaf litter berlese, 22 I 1998, C. Carlton, R. Leschen; 1♂, Hump Ridge, 1220m, 6 II 1976, G.W. Ramsay, litter 76/38; 1♂, Milford rd, 60′, 12 XII 1966, A.K. Walker, K. Wilson, cordyline litter 66/505; 1♂, Eglington Valley Divide, 400m, 13 II 1982, C.F. Butcher, moss/liverworts 82/29; 1♂, Lake Hauroko, 2 II 1966, J.I. Townsend, mixed moss 66/363; 1♀, Beside Tutoko R, 60m, 13 II 1980, J.W. Early, litter, N. menziesii forest; 1♀, Lake Hauroko Road end, 19 II–22 III 2003, FIT, R. Leschen, J. Nunn; 1♂ 1♀ (1♂, slide-mounted), Te Anau Control Gates, #104, 45′26S 167′41E, Nothofagus leaf litter berlese, 22 I 1998, R. Leschen, D. Gleeson; 1♀, Fiordland NP, Kepler tr, above Brod Bay, 680m, 45″24.37′S 167″39.105′E, 11 XII 2005, Nothofagus forest, FMHD #2005- 102, litter, A. Solodovnikov, D. Clarke, ANMT site 1177; 1♀, Fiordland NP, Kepler tr, below Mt. Luxmore Hut, 900m, 45″24.865′S 167″38.675′E, 11 XII 2005, Nothofagus forest, fogging, A. Solodovnikov, D. Clarke, ANMT site 1176; 1♀, Tempest Spur, W. Olivine Ra, 1220–1463m, 25 I 1975, G.W. Ramsay, litter 75/33; Mid Canterbury: 3♂♂, Arthur’ s Pass, Dobson Nature Walk, 8 II 1982, C.F. Butcher, sifted litter 82/26; 3♂♂, Arthur’ s Pass, Bealy tr, 8 II 1982, C.F. Butcher, litter 82/19; 1♂, Craigieburn SF, Dracophyllum Flat tr, 43′10S 171′42E, FIT, #150, 11–27 I 1998, C. Carlton, R. Leschen; North Canterbury: 1♂ 1♀, 2.9km N Arthur’ s Pass, 840m, Bealey Valley Trailhead, 42′55S 171′33E #036, Nothofagus solandri leaf litter berlese, 11 I 1998, C. Carlton, R. Leschen; 1♂, 3km n Arthur’ s Pass, 900m, 42′55S 171′33E, #042, Nothofagus / Dracophyllum leaf litter berlese, 11 I 1998, C. Carlton, R. Leschen; 3♂♂, Arthur’ s Pass NP, Bealey Vy tr, 840m, 18–21 III 1980, subalpine Nothofagus, A. Newton, M. Thayer , litter; 1♂, Arthur’ s Pass NP, Klondyke Cnn, 700m, 26 X 1981, J.W. Early, moss from forest floor; 1♂, Mt. Arthur Pass, Avalanche Peak, 3 I 1950, E. Dawson, leaf litter; 1♂, Arthur’ s Pass NP, Halpine Creek, 17 V 1970, D.S. Homink litter & rotten wood; 1♂, Arthur’ s Pass Mt. Aicken, 950m, 9 II 1984, J.C. Watt, sifted litter 84/24; 1♀, Arthur’ s Pass, Mt. Aicken, 305m, 11 II 1984, J. C. Watt, W. Pond flood debris; Nelson: 5♂♂, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S 172″41.658′E, 29 XI–18 XII 2005, Nothofagus menziesii & N. solandri cliffortioides forest, FMHD #2005-051, FIT, A. Newton, M. Thayer, A. Solodovnikov, AMNT site 1159; 23♂♂ 2♀♀, Kahurangi NP, Cobb Dam rd, Asbestos tr, 450m, 41″06.333′S 172″43.174′E, 29 XI–18 XII 2005, mixed braodleaf (incl. Nothofagus fusca )-podocarp forest, FMHD #2005- 057, carrion(octopus) trap, A. Solodovnikov, D. Clarke, ANMT site 1160; 1♂ 2♀♀, Kahurangi NP, Mt. Arthur tr, below Mt. Arthur Hut, 1200m, 28 XI 2005, Nothofagus forest, FMHD #2005-049, litter, A. Solodovnikov, D. Clarke, ANMT site 1157; 1♀, Cobb Ridge east of Cobb Reservoir, 990m, 2 I 1985, Nothofagus forest, A. Newton, M. Thayer 728, litter; 2♂♂, Kahurnagi NP, Arthur Range, above Flora Saddle, 1000m, 41″11.351′S 172″44.456′E, 28 XI–19 XII 2005, Nothofagus dominant forest, FMHD #2005-044, FIT, A. Newton, M. Thayer, ANMT site 1156; 3♂♂, 0.6km e Gowanbridge, 330m, 18 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 717, FIT &window trap; 3♂♂ 4♀♀, Third House, Dun Mt, 14 IX 1971, G.W. Ramsay, litter 7/114; 1♂ (slide-mounted), 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-podo-nikau forest, A. Newton, M. Thayer 719, window trap; 2♂♂, Karamea sdle, 13 X 1970, J.I. Townsend, litter 70/157; 1♂ 1♀, Karamea Bluff, 1420ft, 29 IX 1966, J. Nunn; 1♂, Cobb Valley, Mytton Hut, 1220m, 13 XII 1967, S. Edridge, litter 67/287; 1♂, Dun Mt, 610m, 4 IV 1966, J.I. Townsend, moss 66/127; 1♂, Arthur Range, w side Flora Saddle, 950m, 1 I 1985, Nothofagus mossy forest, A. Newton, M. Thayer 725, wet litter; 1♂, Punakaiki, Pororari tr, 27 I 1996, forest litter, J. Nunn; 1♂, Oparara Basin, Box Canyon Cave, 200m, 27 II–1 III 2007, J.W. Early, R.F. Gilbert, Nothofagus -podocarp forest, yellow pan trap; 1♂, Glenhope, T. Hall, 30 I 1915; 1♀, Dun Mt, 2000′, 14 II 1942, E.S. Gourlay; 1♀, Dun Mt., 580m, 12 VII 1966, J.I. Townsend, moss 66/204; 1♀, Upper Maitai, 3 V 1950, E.S. Gourlay; 1♀, Wooded Peak, 609m, 31 VIII 1966, J.C. Watt, rotten Nothofagus stump; Otago Lakes: 4♂♂, Mt. Aspiring NP, 12.5km nne Makarora, 370m, 11–17 I 1985, Noth. menz. - hdwd-podocarp forest, A. Newton, M. Thayer 740, FIT &window trap; 1♂ 1♀, Davis Flat, Makarora vly, 19 I 2008, moss, J. Nunn; 1♂, Mt. Aspiring NP, se of Haast Pass, 600m, 11–17 I 1985, Noth. menz. forest, A. Newton, M. Thayer 739, FIT &window trap; 1♂, Mt. Aspiring NP, 5.5km nne Makarora, 330m, 11–17 I 1985, Noth. menz. forest, A. Newton, M. Thayer 741, FIT &window trap; 1♂, Makarora, 23–25 I 1978, 500m, S. & J. Peck, litter; 1♂, McKerrow Range, Makarora, 1150m, 23 I 1978, G. Kuschel, litter 78/51; 1♂, Dart Hut, 13–15 II 1980, J.S. Dugdale, malaise trap; 1♀, 44.5km nw Wanaka, 350m, Matukituki Valley, 44′29S 168′47E, #079, Nothofagus menziesii leaf litter berlese, 18 I 1998, C. Carlton, R. Leschen; 1♀, Cameron Flat, Makarora V, 22 I 1996, J. Nunn; Southland: 1♂, N. Te Anau, 50m, 19 II 1965, A.K. Walker, litter 65/52; 1♂, Owaka Glenomaru Res, 18 I 1978, S. & J. Peck, litter; 1♀, MacLennan, 100’, 14 II 1968, J.I. Townsend, moss 68/20; Westland: 1♂♂ 3♀♀, Okuku Scenic Reserve, 9.2km sse Kumara, 120m, 8–19 I 1986, podocarp-hdwd forest, A. Newton, M. Thayer 731, litter; 1♂, Okuku Scenic Reserve, 9.2km sse Kumara, 120m, 8–19 I 1986, podocarp-hdwd forest, A. Newton, M. Thayer 731, fogging; 2♂♂ 1♀, 1.8km n Punakaiki, 80m, 19 XII 1984 – 20 I 1985, hdwd forest with nikau, A. Newton, M. Thayer 718, FIT &window trap; 5♂♂, 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-podo-nikau forest, A. Newton, M. Thayer 719, window trap; 1♂, 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-podo-nikau forest, A. Newton, M. Thayer 719, litter; 2♂♂, 1.5km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-nikau forest, A. Newton, M. Thayer 720, FIT &window trap; 1♂, 1.5km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd-nikau forest, A. Newton, M. Thayer 720, litter; 2♂♂ 1♀, Lake Ianthe, 7 II 1984, J.C. Watt, sifted litter & wood mould 84/22; 4♂♂ 4♀♀, Hokitika Gorge, 29 I 1978, S.B. Peck, litter; 2♂♂, Okuku State Forest, 10km e of Kumara, 28 VIII 1977, R.M. Emberson, podocarp-broadleaf forest, moss&litter; 1♂, Hokitika R.Gorge, S Kowhitirangi, 100m, 17 III 1980, podocarpbroadleaf, A. Newton, M. Thayer, litter; 1♂ 1♀, Okuku Ck, 11.3km sse Kumara, 60m, 18–22 III 1980, podocarp-broadlf, A. Newton, M. Thayer, litter; 1♂, L. Mahinapua Scen Res, 30m, 16–22 III 1980, podocarp mixed broadleaf, A. Newton, M. Thayer, litter; 1♂ 1♀, Lake Wahapo, 27 XI 1966, J.I. Townsend, litter 66/345; 1♂, Jacksons Bay, 23 IX 1979, A.K. Walker, moss & dead wood litter; 1♂ 1♀, Hokitika Gorge Res, Hokitika, 29 I 1978, S. & J. Peck, litter, 100m; 1♂, Westland NP, Castle Rocks Hut, 1220m, 14–16 I 1996, yellow pan trap, subalpine tussock herb field; 1♂, Haast Pass, road summit, 6 II 2009, sifted ground moss, J. Nunn; 1♂, Haast Pass, 31 I 2005, J. Nunn, forest floor moss; 1♂, Otira, Barretts Creek, 9 II 1982, C.F. Butcher, litter 82/27; 1♂ 1♀, Fantail Falls, 3km ne Haast Pass, 4 II 1984, J.C. Watt, wood mould & litter; 1♂, Summit Haast Pass rd, 1850’, 7 XII 1966, A.K. Walker, litter 66/471; 1♂, Pakihi Walk, 5.2km se Okarito, #062, 43′15S 170′12E, 14 I 1998, litter, C. Carlton, R. Leschen; 1♂, 3.2km ne Haast, 14m, Haast River Walk, 43′52S 169′03E, #072, Nothofagus -podocarp forest litter, 17 I 1998, C. Carlton, R. Leschen; 1♂, 56.6km se Haast, Fantail Falls tr, 475m, 44′05S 169′23E, #073, Nothofagus menziesii litter, 17 I 1998, C. Carlton, R. Leschen; 1♂, Victoria Range, 300’, 26 X 1940, E.S. Gourlay; 3♀♀, Jacksons, Taramakau R, 150m, 26 I 1978, G. Kuschel, sifted litter & rotten wood; 1♀, Haast Pass w of summit, 550m, 24 I 1978, G. Kuschel, sifted litter & rotten wood 78/52; 1♀, Haast Pass, 20 III 1968, R.A. Cumber, litter; 1♀, Otira Valley, Dobsons tr, 1005m, 11 XI 1966, A.K. Walker, moss 66/382; 1♀, Nile River Valley, moss, 1 IX 1971, R.M. Emberson; 1♀, Haast Pass, 518m, 23 X 1966, J.I. Townsend, moss 66/349; 1♀, Lake Kaniere rd, 2.8km nw Lake Kaniere, 120m, 8-19 I 1985, podocarp-hdwd forest, A. Newton, M. Thayer 732, litter; 1♀, 3km n of Bruce Bay, 6 II 2009, sifted moss from podocarp swamp forest, J. Nunn; 1♀, Kelly’ s Ck, Otira, 25 II 1989, J. Nunn.

Diagnosis. This species can be distinguished from other species by the following combination of characters: body larger, length 2.2–2.8 mm; frontal rostrum prominent; anterior and posterior frontal foveae deep and largely oval; ventral surface of male head with distinct triangular process bearing dense setae behind mouthparts; shape of antennomeres and genitalia unique to this species.

Redescription. Length 2.2–2.8 mm. Body reddish to blackish brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 43a ). Head: Round, widest across eyes ( Fig. 43c ). Ventral surface of male head with distinct triangular process bearing dense setae behind mouthparts ( Fig. 43d ). Antennomere 1 approximately 2 times longer than wide, 2 longer than wide, 3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal rostrum prominent. Frontal sulcus deep, slightly exceeding eye ( Fig. 43c ). Anterior and posterior frontal foveae oval ( Fig. 43c ). Eye prominent, one-third length of temple ( Fig. 43c ). Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular ( Fig. 43a ). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe elongate and divided vertically, dorsal process small and triangular ( Fig. 43b ). Phallobase of median lobe symmetrical and rounded ( Fig. 43b ). Parameres symmetrical and wide with setae anteriorly ( Fig. 43b ).

Type Locality. Moa Hill , Mid Canterbury, New Zealand .

Distribution. Buller, Mid Canterbury, North Canterbury, Nelson, Otago Lakes, Southland, Westland ( Fig. 44 : black circles).

Habitat. Most specimens of this species were collected using flight intercept and window traps or by sifting mossy leaf or log litter in hardwood, podocarp, broadleaf, or Nothofagus forests.

Figures

Fig. 43. Sagola strialis (strialis species-group). a) Habitus, dorsal view, b) Aedeagus, dorsal view, c) Dorsal surface of male head, d) Ventral surface of male head. Scale bar a = 1 mm, b–d = 0.1 mm.

Fig. 44. Known collection localities of New Zealand Sagola strialis (circles).