Sagola pertinax Broun, 1893

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 135

Sagola pertinax Broun, 1893

( Figs. 68g , 69g , 70g , 72 )

Sagola pertinax Broun, 1893a: 176 . Raffray 1904: 497, 1911: 5, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.

Type Material. Lectotype. NEW ZEALAND: Waikato: 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “2720. ♂.” [white label, handwritten]; “ Waikato ” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola pertinax ” [white label, handwritten]; “ LECTOTYPE Sagola pertinax Broun Desig. Park and Carlton 2013 ” [red label, printed] . Paralectotypes (n = 2: 1 male; 1 female). NEW ZEALAND: Waikato : 1♂ ( BMNH), glued on rectangular card, “2720. ♂ ” [white label, handwritten]; “ Waikato ” [white label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola pertinax ” [white label, handwritten] . 1♀ ( BMNH), glued on rectangular card, “2720.♀” [white label, handwritten]; “head. elytra” [white label, handwritten]; “Ohaupo. Waikato ” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that three specimens were examined ( Broun, 1893a: 176). This designation will fix the identity of the species and facilitate its recognition by future workers .

Additional Material (n = 3 males). NEW ZEALAND: Auckland : 1♂, Hunua, Clevedon , T. Broun collection ; 1♂, Rangitoto I, 8 XII 1951, moss ; 1♂, Eaves Bush, Orewa , 15 II 1945, A.E. Brookes collection .

Diagnosis. This species can be distinguished from the other species of the insignis species-group by the following combination of characters: body length 2.2–2.5 mm; temporal lobe of male head weakly projecting, as long as eye with flattened ventral surface; ventral surface of male head with triangular setose process behind mouthparts; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.

Redescription. Length 2.2–2.5 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68g ). Head: Bluntly triangular, widest across eyes ( Fig. 69g ). Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Temporal lobe of male head weakly projecting, as long as eye with flattened ventral surface ( Fig. 69g ). Ventral surface of male head with triangular setose process behind mouthparts ( Fig. 69g ). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular ( Fig. 68g ), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, in female as long as wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia broad with narrower rectangular apical lobe ( Fig. 70g ). Phallobase of median lobe symmetrical and rounded ( Fig. 70g ). Parameres symmetrical and slender, with sparse setae from apex to middle ( Fig. 70g ).

Type Locality. Ohaupo, near Mr. Kusab’ s sawmill, Waikato, New Zealand .

Distribution. Auckland, Waikato ( Fig. 72 : black circles).

Habitat. Unknown.

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 72. Known collection localities of New Zealand Sagola species. S. pertinax: black circles; S. monticola: triangles; S. laticeps: black squares; S. castanea: stars; S. sheldoni: white circles; S. mercuryensis: white squares.