Kamopanorpa rasnitsyni, BASHKUEV & SUKATSHEVA, 2021

BASHKUEV, ALEXEY S. & SUKATSHEVA, IRINA D., 2021, New species of Kamopanorpa Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera), Palaeoentomology 4 (5), pp. 453-461 : 454-456

Kamopanorpa rasnitsyni sp. nov.

( Figs 1A, B , 2A )

Type material. Holotype PIN 5386 /9±, part and counterpart (from this point on indicated by ± symbol) of forewing with anal area missing; paratype PIN 5386 /3±, nearly complete forewing of somewhat smaller size.

Etymology. Named in honour of Alex Rasnitsyn, one of the world’s foremost palaeoentomologists, on the occasion of his 85 th birthday.

Diagnosis. SC not reaching fork on RS 1, ending with apical fork, with several (2–4) subcostal crossveins; subcostal space narrow with no visible crossveins.

Locality and horizon. Izykh Coal Mines, quarry no. 1, south of Belyi Yar, Khakassia, Minusinsk Coal Basin, South Siberia ; Izykh Formation, Middle Permian .

Description. Wing oblong and narrow (L:W ratio 1:3–1:3.2), apex at RS 3 –RS 4. SC reaching 2/3 of wing length, ending with apical fork, with four short subcostal crossveins (in paratype 5386/3, only two visible, but may be more). Subcostal space relatively narrow (only slightly broader than costal space), with no visible crossveins other than apical one. Pterostigma not preserved. R nearly straight, apically curved, without pterostigmal branches. RS forking well before middle of wing, with six branches reaching wing margin (in paratype, with five, RS 2 simple). Cell DC closed by crossvein rs 2 –rs 3 located at level of RS 1 fork. Cell MC open. M in holotype with seven branches,

Kamopanorpa lata Martynov, 1928 = Microptysmodes uralicus O. Martynova, 1958

K. rotundipennis Martynov, 1933

K. uralensis (Martynov, 1940)

= Parachorista uralensis Martynov, 1940

= K. fasciipennis O. Martynova, 1942

= K. pallida Sukatsheva, 1976

= K. incerta Sukatsheva, 1976

= Uraloptysma maculata Ivanov, 1992

K. sibirica (O. Martynova, 1958)

= Microptysma sibiricum O. Martynova, 1958

K. pritykinae ( Sukatsheva, 1976)

= Microptysmodes pritykinae Sukatsheva, 1976

K. mongolica Sukatsheva, 1992

= Microptysmodes mongolicus Sukatsheva, 1992

K. sukatchevae Novokshonov, 1993

K. karaungirica Novokshonov, 1993

K. latipennata Novokshonov, 1993

K. grossa Novokshonov, 1993

K. maculosa nom. nov.

= K. maculata Sukatsheva & Aristov, 2013

K. tshepanikhensis Sukatsheva & Aristov, 2013

K. tamarae Sukatsheva, 2017

K. tyulkiana Sukatsheva, 2017

K. rasnitsyni sp. nov.

K. sivchikovi sp. nov.

Middle Permian, Kazanian European Russia, Tatarstan (Tikhie Gory)

Middle Permian, Kazanian European Russia, Arkhangelsk District (Soyana)

Lower Permian, Kungurian European Russia, Cis-Urals (Tshekarda)

Middle Permian, Kazanian, Kuznetsk Fm South Siberia, Kuznetsk Basin (Kaltan, Sarbala-2)

? Upper Permian , Akkolka Fm East Kazakhstan (Karaungir)

Middle Permian Mongolia (Bor-Tologoy)

Middle Permian Mongolia (Bor-Tologoy)

? Upper Permian , Akkolka Fm East Kazakhstan (Karaungir)

? Upper Permian , Akkolka Fm East Kazakhstan (Karaungir)

Middle Permian, Urzhumian European Russia, Udmurtia (Kostovaty = Galevo)

Middle Permian, Urzhumian European Russia, Udmurtia (Tshepanikha; Kostovaty)

Middle Permian, Urzhumian European Russia, Udmurtia (Tshepanikha)

Lower Permian, Ufimian European Russia, Cis-Urals (Tyulkino)

Lower Permian, Ufimian European Russia, Cis-Urals (Tyulkino)

Middle Permian,?Kazanian, Izykh Fm South Siberia, Minusinsk Basin (Belyi Yar)

Middle Permian,?Kazanian, Izykh Fm South Siberia, Minusinsk Basin (Belyi Yar)

K. rotunda sp. nov. Middle Permian,?Kazanian, Izykh Fm South Siberia, Minusinsk Basin (Belyi Yar)

M 1a and M 2 with additional short forks, MP 4 weak, ending blindly in membrane soon after M 3+4 forking point. In paratype, M five-branched. Base of CuA long, strongly oblique, M 5 much shorter. Anal area (preserved only in paratype) with double-Y loop typical for the genus; free distal portion of A 2 very faint. Crossveins mostly faint, visible are: two crossveins in radial space (in holotype, r–rs 1 and r–rs 1a; in paratype, only the distal one); few crossveins between RS branches; two crossveins in RS–M space; long m 3+4 –cua; strong transverse cua–cup located slightly proximal to apex of A 2; and a 1 –a 2 .

Measurements. Holotype: forewing length, as preserved, 4.5 mm, estimated, ~ 5 mm, width, 1.7 mm. Paratype: forewing length, 4 mm, width, 1.3 mm.

Remarks. The new species, as well as the two others described below, differs from the geographically and stratigraphically closest species K. sibirica (O. Martynova, 1958) mainly in much more proximal position of all main forks of RS and M, and possibly in having more subcostal crossveins, although the latter requires further confirmation. Unlike the previous reconstructions ( Martynova, 1958; Novokshonov, 1993b), a re-examination of the holotype of K. sibirica revealed more than one crossvein but the exact number of them is unclear due to poor preservation.

Figures

FIGURE 1. A, B, Kamopanorpa rasnitsyni sp. nov. A, Holotype PIN 5386/9. B, Paratype PIN 5386/3. C, K. rotunda sp. nov., holotype PIN 5386/5. D, K. sivchikovi sp. nov., holotype PIN 5386/1. E, F, Kamopanorpa spp. E, 5386/8. F, 5386/11.All to the same scale.

FIGURE 2. A, Kamopanorpa rasnitsyni sp. nov. Holotype PIN 5386/9. B, K. sivchikovi sp. nov., holotype PIN 5386/1. C, K. rotunda sp. nov., holotype PIN 5386/5. SEM images.