Crusopimpla Kopylov, Spasojevic & Klopfstein, 2018

Kopylov, D. S., Spasojevic, T. & Klopfstein, S., 2018, New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East, Zootaxa 4442 (2), pp. 319-330 : 323

Genus Crusopimpla Kopylov, Spasojevic & Klopfstein , gen. nov.

Type species. Crusopimpla tadushensis Kopylov, Spasojevic & Klopfstein , sp. nov.

Composition. Crusopimpla tadushensis, Crusopimpla ? rediviva Brues.

Diagnosis. Propodeal carinae well developed, enclosing basal area, areola, petiolar area, and first, second and third lateral areas. Fore wing with pterostigma rather wide, about 3× as long as wide; areolet tetragonal with uneven sides, r-m and 2+3-M long, 4-M short and 3Rs absent; ramulus short or absent. Metasomal tergite 1 wider than long, with strong dorsal carinae converging on basal and almost parallel on apical half. Metasomal tergites 2 and following wider than long. Ovipositor robust, about 0.3–0.8 times as long as metasoma. Hind femur thickened.

Etymology. From the Latin word “crus”, which means stem, and the genus name Pimpla , to reflect its basal position within the subfamily.

Remarks. The genus Crusopimpla is similar in general appearance to several extant genera in the subfamily Pimplinae , especially among the Gregopimpla , Pimpla and Scambus genus groups. However, the propodeal carination is always reduced in these groups ( Fig. 1 ). The only extant genera in the subfamily which have a nearly full set of propodeal carinae at least in a few species are Theronia and Xanthopimpla from the tribe Pimplini . They both have a much more elongate pterostigma and radial cell in the fore wing and the ground color of the body is orange or yellow, while C. tadushensis is dark-colored, and basal area of propodeum not developed. The extinct genus Lithoserix Brown also has a similar body shape and the state of its propodeal carinae is somewhat uncertain from the single known fossil; however, Lithoserix also has an elongate pterostigma and radial cell, and possesses a long ramulus in the discocubital cell ( Brown 1986; Kasparyan & Rasnitsyn 1992).

Figures

FIGURE 1. Phylogeny of Pimplinae and outgroups, redrawn according to Gauld et al. (2002), with examples of propodea, first metasomal tergites and hind wings shown to the left of the genus names. The drawings are modified from Townes (1969), and the species representatives are, from top to bottom: Xorides irrigator Fabricius, Odontocolon ochropus Townes, Poemenia hectica Gravenhorst, Rhyssa persuasoria Linnaeus, Theronia atalantae Poda, Lissopimpla excelsa Costa, Xanthopimpla hova Saussure, Echthromorpha agrestoria Swederus, Itoplectis maculator Fabricius, Pimpla madecassa Saussure, Zatypota percontatoria Müller, Dreisbachia mira Tosquinet, Camptotypus pulchripennis Saussure, Scambus tecumseh Viereck, Flavopimpla nava Tosquinet, Delomerista novita Cresson, Atractogaster semisculptus Kriechbaumer, and Perithous albicinctus Gravenhorst.