Crusopimpla Kopylov, Spasojevic & Klopfstein

Kopylov, D. S., Spasojevic, T. & Klopfstein, S., 2018, New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East, Zootaxa 4442 (2), pp. 319-330: 323

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Crusopimpla Kopylov, Spasojevic & Klopfstein

gen. nov.

Genus Crusopimpla Kopylov, Spasojevic & Klopfstein  , gen. nov.

Type species. Crusopimpla tadushensis Kopylov, Spasojevic & Klopfstein  , sp. nov.

Composition. Crusopimpla tadushensis, Crusopimpla  ? rediviva Brues. 

Diagnosis. Propodeal carinae well developed, enclosing basal area, areola, petiolar area, and first, second and third lateral areas. Fore wing with pterostigma rather wide, about 3× as long as wide; areolet tetragonal with uneven sides, r-m and 2+3-M long, 4-M short and 3Rs absent; ramulus short or absent. Metasomal tergite 1 wider than long, with strong dorsal carinae converging on basal and almost parallel on apical half. Metasomal tergites 2 and following wider than long. Ovipositor robust, about 0.3–0.8 times as long as metasoma. Hind femur thickened.

Etymology. From the Latin word “crus”, which means stem, and the genus name Pimpla  , to reflect its basal position within the subfamily.

Remarks. The genus Crusopimpla is similar in general appearance to several extant genera in the subfamily Pimplinae  , especially among the Gregopimpla  , Pimpla  and Scambus  genus groups. However, the propodeal carination is always reduced in these groups ( Fig. 1 View Figure ). The only extant genera in the subfamily which have a nearly full set of propodeal carinae at least in a few species are Theronia  and Xanthopimpla  from the tribe Pimplini  . They both have a much more elongate pterostigma and radial cell in the fore wing and the ground color of the body is orange or yellow, while C. tadushensis  is dark-colored, and basal area of propodeum not developed. The extinct genus Lithoserix Brown  also has a similar body shape and the state of its propodeal carinae is somewhat uncertain from the single known fossil; however, Lithoserix  also has an elongate pterostigma and radial cell, and possesses a long ramulus in the discocubital cell ( Brown 1986; Kasparyan & Rasnitsyn 1992).