Manota biunculata, Hippa, Heikki, 2007

Hippa, Heikki, 2007, The genus Manota Williston (Diptera: Mycetophilidae) in Melanesia and Oceania, Zootaxa 1502, pp. 1-44: 25-27

publication ID 10.5281/zenodo.177118

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scientific name

Manota biunculata

sp. n.

Manota biunculata  sp. n.

( Figs 13View FIGURE 13 A, B)

A small-sized Manota  .

Male. Coloration. Pale yellowish-brown, frons, vertex and occiput darker, mouthparts and legs slightly paler than thorax, wing unicolorous, hyaline, pale yellowish-brown, haltere yellowish-brown. Head. Antennal flagellomere 4, in the single specimen, seen in oblique ventral view and not illustrated, the lateral view apparently similar to Fig. 10View FIGURE 10 C. Palpomere 3 of maxillary palpus with apicomesial extension, with 5 apically expanded curved sensilla; palpomere 4 with parasegment. Thorax. Anepisternum setose, with 37 setae; anterior basalare non-setose; preepisternum 2 setose, with 9 setae; laterotergite setose, with 9 setae; episternum 3 setose, with 8 setae. Wing. Length 1.4 mm. Hypopygium ( Figs 13View FIGURE 13 A, B). Sternite 9 extending halfway between the base of gonocoxa and the base of gonostylus, laterally sharply delimited, posterior margin transversely straight, anterior margin deeply incised, setae similar to ventral setae of gonocoxa. Parastylar lobe exposed, with 1 transverse seta arising from a small lobe on its anterior part. Paraapodemal lobe exposed in ventral view. The dorsal mesial margin of gonocoxa simple, subapically with an aggregation of setae, mostly on ventral side. Two long apically curved juxtagonostylar megasetae present, arising from a large basal body, which is about half the length of megasetae. Gonostylus elongate, with long setae, dorsally also with short setae some of which are on a low subapical transverse crest. Tegmen broadly triangular, with weakly expressed shoulders. Hypoproct small, posteriorly extending to the level of the middle of gonocoxa, with ca. 7 ventral setae on each side in a rather tight anterolateral group. Cerci separate.

Female and preimaginal stages unknown.

Discussion. M. biunculata  resembles M. gemella  and M. tricuspis  . It is distinguished from both by having two subequal juxtagonostylar megasetae, not one megaseta and a shorter normal seta, and by lacking a long apical lobe dorsally on gonocoxa, which is apicomesial in M. tricuspis  and apicolateral in M. gemella  . Furthermore, it differs from M. tricuspis  by lacking the three huge setae apically on gonostylus and from M. gemella  by lacking a small lobe in the middle of the mesial margin of gonostylus. M. biunculata  is also similar to the Oriental M. orientalis Senior-White  but differs by the same characters as M. gemella  . M. biunculata  resembles the Oriental M. angustata Hippa  but differs e.g. by having the gonostylus much broader, 1 / 3 of the length instead of 1 / 5, by lacking a distinct subapical mesial lobe on the gonostylus, and by having the two juxtagonostylar megasetae subequal in size, not conspicuously unequal. M. biunculata  also resembles the Oriental M. ovata Hippa  but differs e.g. by having the mesial margin of gonostylus concave, not convex, by lacking a conspicuous fringe of long setae on this margin, and by lacking a distinct transverse lobe dorso-apically on the gonocoxa. The hypopygium of M. biunculata  is not dissimilar to the Oriental M. calcarata Hippa  , M. procera Hippa  and M. transversa Hippa  , but all these species have a long apicolateral lobe dorsally on the gonocoxa and a non-setose laterotergite.

Types. Holotype. Male, Papua New Guinea, Madang prov., Halopa village, primary rainforest, 600 m a. s. l., Malaise trap, November 2000, Lukáš Č iżek leg. (in NRM).


Swedish Museum of Natural History - Zoological Collections