Alloxysta victrix ( Westwood, 1833 )

Ferrer-Suay, Mar, Selfa, Jesús & Pujade-Villar, Juli, 2018, Palaearctic species of Charipinae (Hymenoptera, Figitidae): two new species, synthesis and identification key, European Journal of Taxonomy 427, pp. 1-110 : 76-79

Alloxysta victrix ( Westwood, 1833)

Figs 3.17, 5.21

Allotria victrix Westwood, 1833: 495 . Type: deposited in OUMNH ( Andrews, 1978: 92) (not examined).

Xystus victrix – Rondani 1878: 177.

Allotria (Allotria) victrix – Dalla Torre & Kieffer 1902: 41.

Charips (Charips) victrix victrix – Dalla Torre & Kieffer 1910: 285.

Charips victrix – Rohwer & Fagan 1917: 360.

Alloxysta victrix – Hellén 1963: 16.

Alloxysta victrix victrix – Andrews 1978: 92.

Diagnosis

Alloxysta victrix is mainly characterized by a large closed radial cell that is 3.0 times as long as wide (Fig. 5.21); the presence of pronotal carinae; the absence of propodeal carinae; the lack of setae on longitudinal areas where carinae are present in other species of Charipinae (Fig. 9.2); and male and female antenna with rhinaria beginning on F3, F1 longer than pedicel and F2, F2–F4 subequal (Fig. 3.17), and F1–F3 curved in males. It is similar to A. consobrina , but can be differentiated by the proportions of the flagellomere (F2–F4 are subequal in length in A. victrix (Fig. 3.17), while F2 is subequal to F3 and F3 is shorter than F 4 in A. consobrina (Fig. 2.14)), the size of the radial cell (3.0 times as long as wide in A. victrix (Fig. 5.21), but 2.7 times in A. consobrina (Fig. 4.12)), and the degree of propodeal pubescence (in A. victrix , the propodeum lacks setae at the longitudinal area where the carinae are present in other Charipinae , while in A. consobrina , the propodeum is completely covered with dense setae).

Material examined (376 ♂♂, 522 ♀♀)

CYPRUS: 1 ♀ [Limassol, Feb. 1934, G.A. Mavromoustakis] (BMNH B.M. 1935-55); 1 ♂, 2 ♀♀ [Limassol, 25 May 1934, G.A. Mavromoustakis] (BMNH B.M. 1935-55); 1 ♀ [Kilani, Krios R., 11 Sep. 1937, G.A. Mavromoustakis] (BMNH B.M. 1937-808).

CZECH REPUBLIC: 1 ♀ [Bohemia: Praha-Kunratice, 18 Jun. 1964, P. Mikula] (BMNH); 6 ♀♀ [Moravia, Dyje River, near Znojmo, 12 Aug. 1991, L. Masner] (CNCI C-263); 1 ♀ [central Bohemia, 30 Jul. 1991, J. Macek] (CNCI C-274); 15 ♂♂, 39 ♀♀ [Moravia, Palava near Mikulov, = 400 m, 9 Aug. 1991, L. Masner, sweep] (CNCI C-264); 9 ♂♂, 3 ♀♀ [Moravia, Lanzhot-Ranspurk, 7–9 Aug. 1991, L. Masner, climax flood forest] (CNCI C-265); 2 ♂♂, 10 ♀♀ [Moravia, Tvrdonice, 7 Aug. 1991, L. Masner, fraxinetum] (CNCI C-262); 11 ♂♂, 15 ♀♀ [Moravia, Lanzhot-Ranspurk, 7–9 Aug. 1991, L. Masner, sweep, climax flood forest] (CNCI C-268); 2 ♂♂, 9 ♀♀ [Moravia, 16 km N Blansko, 8 Aug. 1991, L. Masner, sweep, Tilia-Acer forest] (CNCI C-271); 18 ♂♂, 14 ♀♀ [Moravia, Lanzhot-Ranspurk, 7–9 Aug. 1991, L. Masner, sweep, climax flood forest] (CNCI C-269); 1 ♂, 7 ♀♀ [Moravia, Javorina, 800–900 m, 10 Aug. 1991, L. Masner, sweep, climax deciduous forest] (CNCI C-266); 2 ♂♂, 2 ♀♀ [Moravia, Lanzhot-Ranspurk, 9–12 Aug. 1991, L. Masner, climax hardwood forest, PT] (CNCI C-270); 1 ♀ [Moravia, Vranov, Dyje River, 13 Aug. 1991, L. Masner] (CNCI C-273); 13 ♂♂, 26 ♀♀ [Moravia, Ladnice environs 7–9 Aug. 1991, L. Masner, riparian forest] (CNCI C-272); 1 ♀ [Praha-Troja, 6–9 Sep. 1999, YPT, L. Masner, steppe] (CNCI C-284); 2 ♀♀ [Moravia, Zidlochovice, 20 Nov. 1984, N.D. Springate] (BMNH).

DENMARK: 1 ♀ [W. Zealand, Gyrstinge, 20 km S of Holbaek, 28 Aug. 1994] (CNCI C-243).

FRANCE: 2 ♀♀ [Montpelier, 1980 Dufom (handwritten), Alloxysta victrix (Westwood) ♀ det. J. Quinlan, 1987] (BMNH); 8 ♂♂, 10 ♀♀ [Pyrénées Orientalis nr. Arñes-sur. Tech, D. and J. Clark, 23 May 1961] (BMNH B.M. 1962-149); 2 ♂♂, 3 ♀♀ [St. Genis-Loval, Jun. 1928, Ex. Macrosiphum dirhodum Walk, R. Pussard , Charips victrix Wstw , ♀ Ch. Ferriere det] (BMNH); 1 ♀ [St. Genis-Laval, Jun. 1928, Ex Macrosiphum dirhodum, Walk., R. Pussard , Charips victrix Westw., Ch. Ferriere det., Pres. by Imp. But. Ent.] (BMNH B.M. 1929-196); 1 ♀ [B. du Rhone Fonscolombe (3), 2 Jul. 1980, M.W.R. de V. Graham coll.] (BMNH B.M. 1995-489); 1 ♀ [Hérault, Baillarguet CSIRO lab., 43°41′12″ N, 3°52′24″ E, 15–22 May 1993, P.G. Mason, champ sauvage, MT] (CNCI C-302); 2 ♀♀ [Haute Savoie, Bossy sur Frangy, 11–19 Sep. 1993, J. Steffen, on wood pile] (CNCI C-188); 1 ♀ [Hérault, Baillarguet, 43°42′18″ N, 3°53′ E, 12–24 Jun. 1995, P.G. Mason, pine look forest, PT and MT] (CNCI C-301); 2 ♀♀ [Haute Savoie, Bossy sur Frangy, 17–30 Aug. 1993, J. Steffen, on wood pile] (CNCI C-289); 1 ♀ [Hérault, Montferrier sur Lez, 43°40′ N, 3°51′ E, 23 May 1994, P.G. Mason, YPT] (CNCI C-300).

GERMANY: 1 ♂ [Baden Woldkrich, A. Koebele] (BMNH B.M. 1942-95); 3 ♂♂, 3 ♀♀ [Stuttgart, 21 Jul. 1937, T.H. Rowsell and B.J. Clifton] (BMNH B.M. 1937-539); 1 ♂, 2 ♀♀ [Wern, Riedenerwald, 15 Aug. 1978, L. Huggert, sweep, in open oak wood] (CNCI C-234); 1 ♀ [Mainz, 18 Sep.–1 Oct. 1965, A.W. Steffan] (CNCI C-236); 1 ♀ [Munich, 23 Jul. 1984, R. Wharton] (CNCI C-239); 1 ♀ [Mainz, 26 Aug.–3 Sep. 1965, A.W. Steffan] (CNCI C-231); 2 ♀♀ [Mainz, 18–28 Jun. 1968, A.W. Steffan] (CNCI C-226); 1 ♀ [Ingelheim am Rhein, MT, 1 May–5 Jun. 1969, A.W. Steffan] (CNCI C-227); 1 ♂, 4 ♀♀ [Ingelheim am Rhein, ??.??.1968, MT in orchard, A.W. Steffan] (CNCI C-238); 6 ♀♀ [Ingelheim am Rhein, 1–30 Sep. 1968, MT, A.W. Steffan, orchard, Cynipoidea ] (CNCI C-237); 2 ♀♀ [Mainz, 15 Nov. 1965, A.W. Steffan] (CNCI C-233); 15 ♂♂, 82 ♀♀ [Schwarzwald, Todtmoos environs, 6 Aug. 1984, L. Masner, screen sweeping] (CNCI C-235); 33 ♂♂, 20 ♀♀ [Anmuhle, near Hamburg Sachsenwald, 22 Aug. 1984, L. Masner, screen sweeping] (CNCI C-296).

IRAN: 1 ♀ [Karaj, 40 km W Tehran, 25–28 Jun. 1978, J.T. Huber, YPT] (CNCI C-316).

ITALY: 1 ♂ [Cortina D’Ampezzo, 24 Jul. 1969, M.C. Day Swept in grall] (BMNH); 1 ♂ [Orta, 5 Apr. 1950, G.J. Kerrich] (BMNH); 1 ♀ [Dolomites, Seis am Schlein, 1–13 Jun. 1964] (BMNH); 3 ♀♀ [Abruzzo (AQ), L’Aquila, Aterno River, sweeping riparian veg. and Salix , 20 Jun. 1992, J. D. Pinto] (CNCI C-253).

JAPAN: 1 ♂ [Hokkaido, Hidako Mts below Pyo Tan, 500 m, 14 Aug. 1996, L. Masner, S.S. J-12] (CNCI C-161); 1 ♀ [Tsukuba, Ibaraki, 21 Jun. 1987, sweep, C.M. Yoshimoto] (CNCI C-182); 1 ♀ [Hokkaido, Furano, Exp. Forest, 43°15′ N, 142°20′ E, 9 Aug. 1996, 500 m, L. Masner, sweep] (CNCI C-163); 1 ♀ [Ibaraki, Tsukuba, Expo site, 21–31 May 1989, M.J. Sharkey] (CNCI C-214).

F.Y.R MACEDONIA: 1 ♀ [Verge of oak wood above lake, Yugoslavia, Macedonia, Prespa Geril, 22 Jun. 1958, R.L. Coe.] (BMNH B.M. 1958-417).

MALTA: 3 ♂♂, 1 ♀ [May 1982, sp C.I.E. A. 13986, Alloxysta Först ? sp. ♀ det. J. Quinlan, 1982] (BMNH); 1 ♂, 1 ♀ [Malta, May 1982, sp. DI, C.I.E., A. 13986] (BMNH).

MOROCCO: 1 ♀ [Middle Atlas, flowers at edge of wood, Frane 5.500 ft, 10 Jun. 1961, P.N. Lawrence] (BMNH B.M. 1961-328); 18 ♀♀ [Marrakech, Ouigane, 1000 m, 31°08′ N, 8°05′ W, 3–8 May 1996, MT, C. Kassebeer] (CNCI C-291); 6 ♀♀ [Marrakech, Ouirgane, 1000 m, 31°08′ N, 8°05′ W, 8–13 May 1996, C. Kassebeer, MT] (CNCI C-295); 2 ♀♀ [Marrakech, Ouirgane, 1000 m, 31°08′ N, 8°05′ W, 4–11 Oct. 1996, C. Kassebeer, MT] (CNCI C-279); 1 ♀ [Marrakech, Ouirgane, 1000 m, 31°08′ N, 8°05′ W, 19–25 Mar. 1997, C. Kassebeer, MT] (CNCI C-281); 1 ♀ [Marrakech, Ouirgane, 1000 m, 31°08′ N, 8°05′ W, 11–17 Jun. 1996, C. Kassebeer, MT] (CNCI C-282); 1 ♀ [Marrakech, Ouirgane, 1000 m, 31.08′ N, 8.05′ W, 17–23 Jun. 1996, C. Kassebeeer] (CNCI C-283).

NETHERLANDS: 1 ♂, 3 ♀♀ [Wageningen, 26 Jul. 1978, L. Huggert, swept in marsh] (CNCI C-310).

POLAND: 1 ♂, 2 ♀♀ [Przemysl, 12 Sep. 1984, N.D. Springate] (BMNH).

PORTUGAL: 1 ♀ [Gondomar Douto Litoral, 18 Jun. 1962, NLH Krauss] (USNM).

ROMANIA: 1 ♀ [Brasov, 13 Km S. of Brasov, vegetation under prunes, 11–12 Aug. 1967, B.H. and M.C., Cogan] (BMNH B.M. 1969-270).

RUSSIA: 1 ♀ [Russian Far East, Primorski Krai Lazovski Zapovednik, c. 170 km E. of Vladivostok, Ta- Chingousa, 43°00′42″ N, 134°07′34″ E, Om 28 Aug.–16 Sep. 2001, sandy coast, Malaise trap 494°, M. Quest coll., Alloxysta sp4, det. M. Forshage 2012] (BMNH B.M. 2009-59); 1 ♀ [Primoskiy Kray Ussyriysk District, Gomotayozhnoye, 11–15 Aug. 2003, Malaise trap, M.V. Michailovskaya] (USNM).

SLOVAKIA: 1 ♂, 11 ♀♀ [Slovakia, Čachtice environs, 2 Aug. 1991, L. Masner, sweep] (CNCI C-267).

SLOVENIA: 1 ♂ [Postojne, 18 Jul. 1958, R.L. Coe, herbage in mixed forest] (BMNH B.M. 1958-417); 1 ♀ [Postojne, 18 Jul. 1958, R.L. Coe, wooded hill N.W. of town] (BMNH B.M. 1958-417); 1 ♂ [Slatna, Radovljica, 6 Aug. 1978, L. Huggert, swept on open wooded (deciduous) meadow] (CNCI C-275).

SWEDEN: 1 ♂ [Sk. Lund Zoological Mus. grounds Aug. 1976: 6 ♂ and 20 ♀; Sö. Sandemar, T.H. and J.Q., Aug. 1976] (BMNH); 4 ♂♂ [SK. Snogholmss jön, T.H. and J.Q., Aug. 1976] (BMNH); 3 ♂♂, 2 ♀♀ [Sk. Fyladalen, T.H. and J.Q., Aug. 1976] (BMNH); 6 ♀♀ [Sk., Rolsberga, Rovurekulan, RN- 1354/6188] (BMNH); 1 ♀ [Up. Solna Bergshamra, T.H. and J.Q., Aug. 1976] (BMNH); 1 ♀ [Sö, Sandemars slott, T.H. and J.Q., Aug. 1976] (BMNH); 1 ♀ [Up., Salna Bergshamra, T.H. and J.Q., Aug. 1976] (BMNH); 1 ♀ [Sö, Sandemars slott, T.H. and J.Q., Aug. 1976] (BMNH); 1 ♀ [Sö, Dalarö Malmenm, Aug. 1976] (BMNH); 1 ♀ [Sk., Rolsberga Rovurekulan, RN-1354/6188] (BMNH); 2 ♂♂, 13 ♀♀ [Sk., Silvåkra Stensoffa, T.H. and J.Q., Aug. 1976, RN-1351/6176] (BMNH); 1 ♀ [So Malmen, Dalaro, J.Q. and T.H., 15–26 Aug. 1976] (BMNH); 1 ♂ [Sk., Snogholmssjön, T.H. and J.Q., Aug. 1976] (BMNH); 1 ♀ [Sö, Sandemar, T.H. and J.Q., Aug. 1976] (BMNH); 1 ♀ [Sö, Dalarö Malmen, Aug. 1976] (BMNH); 8 ♂♂, 20 ♀♀ [Sk. Höör distr., D.M.S.P. and J.F.P., 8 Jun. 1938] (BMNH B.M. 1938-414); 1 ♀ [Sk, S. Sandby Maryd, T.H. and J.Q., Aug. 1976] (BMNH); 1 ♀ [Sö, Gålö, Stegsholm, T.H. and J.Q., Aug. 1976] (BMNH); 5 ♀♀ [Sk., Lund Zoological Mus. grounds, Aug. 1976] (BMNH); 1 ♀ [Sk., Rolsberga Rovurekulan, T.H. and J.Q., Aug. 1976, RN-1354/6188] (BMNH); 1 ♀ [So, Ludgo s:n, Tovetorp Field Station, 58.947750, 17.148542, sweep, around station rounds, M. Gates and M. Buffington, 6 Aug. 2012] (USNM).

SWITZERLAND: 201 ♂♂, 85 ♀♀ [Dielsdorf, 650 m, 17 Aug. 1984, L. Masner, sweep] (CNCI C-290); 1 ♀ [Solothurn, Weissenstein, 1225 m, 47°15′10″ N, 7°30′00″ E, 17 Jun. 1999, Goulet/White, lush meadow] (CNCI C-285); 3 ♀♀ [Turgau, Unterwasser environs, 1440 m, 4 Aug. 1984, L. Masner, sweeping] (CNCI C-293).

UNITED KINGDOM: England: 4 ♀♀ [Lincolnshire, Moorside near Fulsby Wood, 22–25 Aug. 1986, 53°7′6″ N, 0°7′11″ W, J.S. Noyes] (CNCI C-277); 1 ♂ [Torquay, 25 Jun. 1960, J.R. Vockeroth] (CNCI C-299).

Distribution

Cosmopolitan.

Certain records: Andorra ( Ferrer-Suay et al. 2011: 359), Corsica ( Ferrer-Suay et al. 2013l), England ( Westwood 1833: 495; Müller et al. 1999: 346; van Veen et al. 2003: 450), Japan ( Ferrer-Suay et al. 2013i), Madeira ( Borges et al. 2008; Ferrer-Suay et al. 2012g: 13), Mexico ( Ferrer-Suay et al. 2013m:

37), New Zealand ( Valentine 1975: 59; Ferrer-Suay et al. 2012h: 237), Serbia and Montenegro (Ferrer- Suay et al. 2013b: 363).

Uncertain records: Australia ( Girault 1932: 3), Austria ( Giraud 1860: 127; Hellén 1963: 16), Belgium ( Fabianus 1900), Brazil ( Peronti et al. 2007: 132), Canada ( Fitch 1861: 841), Chile ( Guerra et al. 1998: 335), England ( Curtis 1838: 688; Cameron 1883: 366; Dalla Torre & Kieffer 1910: 285, 288; West et al. 1998: 1458), Finland ( Hellén 1963: 6), France ( Kieffer 1902a: 15–16; 1902b: 70, 600; De Gaulle 1908: 26), Germany ( Hartig 1840: 199; Hübner et al. 2002: 507; Höller et al. 1993: 13), Greenland ( Buhl 1997: 164), Hungary ( Dalla Torre & Kieffer 1910: 285; Fülöp et al. 2010: 55), Ireland ( O’Connor & Nash 1997), Israel ( Argaman 1988: 114), Italy ( Pagliano 1995: 3), Lappland ( Zetterstedt 1838: 410; Hellén 1963: 16), Norway ( Hellén 1966: 393; Westrum et al. 2010), Poland ( Kierych 1979b: 15; Krawczyk et al. 2009: 161), Romania ( Ionescu 1969: 261; Feraru & Mustata 2005: 75), Russia ( Belizin 1962: 127; Hellén 1963: 16; Bokina 1997: 435), Scotland ( Cameron 1883: 366), Spain ( Torras-Casals 1996: 196), Sweden ( Thomson 1862: 406; 1877: 814), The Netherlands ( Hellén 1963: 16; Andrews 1978: 92), USA ( Fitch 1861: 841; Kieffer 1909: 481; Sullivan & van den Bosch 1971; Mertins 1985: 186).

New records: Cyprus, Czech Republic, Denmark, Iran, Malta, Morocco, Portugal, Romania, The Netherlands and Switzerland.