Copuetta, Haddad, Charles R., 2013
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publication ID |
https://doi.org/ 10.11646/zootaxa.3688.1.1 |
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publication LSID |
lsid:zoobank.org:pub:DB84C679-C2AA-41CC-816E-83E2F6DCA391 |
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DOI |
https://doi.org/10.5281/zenodo.6153929 |
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persistent identifier |
https://treatment.plazi.org/id/03DB8445-FF8D-FFA8-FF2A-FBED31C6FF3A |
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treatment provided by |
Plazi |
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scientific name |
Copuetta |
| status |
gen. nov. |
Key to the species of the genus Copuetta View in CoL gen. nov.
1 Males (3 of C. kakamega View in CoL sp. nov. and C. litipo View in CoL sp. nov. unknown).............................................. 2
– Females (Ƥ of C. lesnei View in CoL sp. nov., C. uzungwa View in CoL sp. nov. and C. wagneri View in CoL sp. nov. unknown)........................... 12
2 Embolus with very broad base and nearly transverse fine curved distal section, tip slightly curved and directed prolaterally ( Fig. 104 View FIGURES 99 – 109 )................................................................................ C. lotzi View in CoL sp. nov.
– Embolus with a narrow base, forming at least one complete coil................................................. 3
3 Embolus forming a distinctly compressed coil with one or more spirals (e.g. Figs 99–103 View FIGURES 99 – 109 )............................ 4
– Embolus not forming a spiraling compressed coil, somewhat elongate, with retrolaterally directed tip ( Figs 105, 107, 108 View FIGURES 99 – 109 )...10
4 Embolus short, nearly straight, directed at distal end of cymbium, with three complete coils ( Figs 92 View FIGURES 92 – 98 , 100 View FIGURES 99 – 109 )...................................................................................................... C. erecta View in CoL sp. nov.
– Embolus otherwise, with general orientation of coils and embolus tip oblique, with one or two complete coils............. 5
5 Embolus with length of tip approximately equal to the width of the distal coil ( Figs 106, 109 View FIGURES 99 – 109 )......................... 6
– Embolus with length of tip less than half the width of distal coil................................................. 7
6 Base of embolus on prolateral side slightly narrowed, continuous with tegulum; embolus tip directed 45° prolaterally ( Fig. 106 View FIGURES 99 – 109 ).................................................................................. C. maputa View in CoL sp. nov.
– Base of embolus on prolateral side strongly narrowed, appearing notched; embolus tip directed to distal end of cymbium ( Fig. 109 View FIGURES 99 – 109 )................................................................................. C. wagneri View in CoL sp. nov.
7 Tibiae III with a single ventral terminal spine................................................................ 8
– Tibiae III without a ventral terminal spine.................................................................. 9
8 Embolus with sharp triangular tip directed at distal end of cymbium ( Fig. 103 View FIGURES 99 – 109 ); palpal tegulum with distinct concavity along retrolateral margin, just beyond midpoint between base and tip ( Fig. 128 View FIGURES 128 – 131 ); femora IV with single retrolateral spine on dorsal surface................................................................................. C. lesnei View in CoL sp. nov.
– Embolus with blunt tip directed retrolaterally distally ( Figs 93 View FIGURES 92 – 98 , 102 View FIGURES 99 – 109 ); palpal tegulum without distinct concavity, straight along retrolateral margin beyond midpoint ( Fig. 124 View FIGURES 124 – 127 ); femora IV with two retrolateral spines on dorsal surface................................................................................................. C. lacustris ( Strand, 1916) View in CoL
9 Palpal tegulum tapering gradually to distal embolus ( Fig. 99 View FIGURES 99 – 109 ); embolus with short, relatively broad base, two compressed coils and triangular tip directed distally ( Fig. 99 View FIGURES 99 – 109 ) ( Comoros)........................................ C. comorica View in CoL sp. nov.
– Palpal tegulum with slight bump retrolaterally near embolus base ( Fig. 101 View FIGURES 99 – 109 ); embolus base narrow and stalk-like; embolus tiny, with single compressed coil, tip directed retrolaterally distally ( Fig. 101 View FIGURES 99 – 109 ) ( Tanzania)............ C. kwamgumi View in CoL sp. nov.
10 Embolus long, base directed at distal end of cymbium, narrowly coiled around longitudinal axis with gentle prolateral bend ( Figs 95 View FIGURES 92 – 98 , 105 View FIGURES 99 – 109 ); tibiae I without pair of prolateral spines on their dorsal surface....................... C. magna View in CoL sp. nov.
– Embolus short, base directed prolaterally distally, with sharp retrolateral bend ( Figs 107, 108 View FIGURES 99 – 109 ); tibiae I with pair of prolateral spines on their dorsal surface........................................................................... 11
11 Metatarsi II with single retrolateral spine on their dorsal surface; embolus with narrow base, broadening distally, with very short distally directed retrolateral tip ( Figs 108 View FIGURES 99 – 109 , 149 View FIGURES 149 – 152 .......................................... C. uzungwa View in CoL sp. nov.
– Metatarsi II without retrolateral spine on their dorsal surface; embolus with narrow base, tapering distally, with fine tip directed retrolaterally ( Figs 107 View FIGURES 99 – 109 , 145 View FIGURES 145 – 148 ).......................................................... C. naja View in CoL sp. nov.
12 Epigyne with large peripheral subtriangular ridge, small spermathecae and oblique adjacent median copulatory openings ( Fig. 139 View FIGURES 137 – 140 ).................................................................................. C. magna View in CoL sp. nov.
– Epigyne lacking large peripheral ridge surrounding entire epigyne; spermathecae large; copulatory openings clearly separated and not situated medially............................................................................... 13
13 Epigyne with distinct large sclerotised ridges forming depression including copulatory openings (e.g. Figs 97, 98 View FIGURES 92 – 98 , 116 View FIGURES 114 – 117 ).... 14
– Epigyne without distinct ridges and depression; copulatory openings small, sometimes surrounded by sclerotised rim ( Figs 122 View FIGURES 120 – 123 , 130 View FIGURES 128 – 131 , 135 View FIGURES 133 – 136 , 147 View FIGURES 145 – 148 )................................................................................... 18
14 Entire epigynal ridge situated within the margins of the spermathecae ( Figs 118 View FIGURES 118 – 119 , 126 View FIGURES 124 – 127 ).............................. 15
– Part of epigynal ridges extending beyond margins of spermathecae ( Figs 112 View FIGURES 110 – 113 , 143 View FIGURES 141 – 144 ), otherwise epigynal ridges posteriorly situated and not passing over spermathecae ( Fig. 116 View FIGURES 114 – 117 )........................................................... 16
15 Epigynal ridges large and round; ST II large and round; ST I with many folds ( Fig. 118 View FIGURES 118 – 119 )............. C. kakamega View in CoL sp. nov.
– Epigynal ridges comma-shaped; ST II oval, obliquely orientated; ST I without folds ( Fig. 126 View FIGURES 124 – 127 ).... C. lacustris ( Strand, 1916) View in CoL
16 Epigyne with weakly sclerotised semi-circular ridges, curving laterally and posteriorly, not passing over spermathecae; copulatory openings within small coiled ridges ( Fig. 116 View FIGURES 114 – 117 )............................................. C. erecta View in CoL sp. nov.
– Epigyne with strongly sclerotised semi-circular ridges, curving over posterior half of ST II; copulatory openings round, just behind anterior margin of epigynal ridges ( Figs 98 View FIGURES 92 – 98 , 143 View FIGURES 141 – 144 )..................................................... 17
17 Epigyne with semicircular ridges, meeting medially; ST I and ST II equal in size and width ( Fig. 112 View FIGURES 110 – 113 )....................................................................................................... C. comorica View in CoL sp. nov.
– Epigyne with oval ridges, lateral and anterior margins at nearly right angle; ST II much larger and broader than ST I ( Figs 98 View FIGURES 92 – 98 , 143 View FIGURES 141 – 144 )................................................................................. C. maputa View in CoL sp. nov.
18 Epigyne with strongly recurved copulatory openings directed anteriorly, not surrounded by a sclerotised rim; ST II clearly broader than ST I ( Fig. 135 View FIGURES 133 – 136 ) ( South Africa)..................................................... C. lotzi View in CoL sp. nov.
– Epigyne with round, oval or comma-shaped copulatory openings situated within a sclerotised rim; ST I and ST II equal in width ( Figs 122 View FIGURES 120 – 123 , 130 View FIGURES 128 – 131 , 147 View FIGURES 145 – 148 ) ( Tanzania).................................................................... 19
19 Epigyne with large oval sclerotised rim laterally in posterior half of epigyne, enclosing transverse oval copulatory openings ( Fig. 130 View FIGURES 128 – 131 )............................................................................... C. litipo View in CoL sp. nov.
– Epigyne with small comma-shaped sclerotised rim laterally at midpoint or in anterior half of epigyne, enclosing round or comma-shaped copulatory openings ( Figs 122 View FIGURES 120 – 123 , 147 View FIGURES 145 – 148 )......................................................... 20
20 Epigyne with large indistinct semi-circular ridges at midpoint of epigyne, surrounding large inverted comma-shaped sclerotised rims in anterior half of epigyne; copulatory openings inverted comma-shaped ( Fig. 147 View FIGURES 145 – 148 )............. C. naja View in CoL sp. nov.
– Epigyne without indistinct semi-circular ridges; sclerotised rims small, shaped like transverse commas, at midpoint of epigyne; copulatory openings small and round ( Fig. 122 View FIGURES 120 – 123 )............................................. C. kwamgumi View in CoL sp. nov.
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