Bryocoris, FALLEN, 1829, Fallen, 1829

Konstantinov, Fedor V. & Knyshov, Alexander A., 2015, The tribe Bryocorini (Insecta: Heteroptera: Miridae: Bryocorinae): phylogeny, description of a new genus, and adaptive radiation on ferns, Zoological Journal of the Linnean Society 175 (3), pp. 441-472 : 460

publication ID 10.1111/zoj.12283


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FIGS 4, 5 View Figures 1–9 , 14, 20 View Figures 10–21 , 23, 29 View Figures 22–33 , 36 View Figures 34–45 , 57 View Figures 46–58 , 62, 68, 69 View Figures 59–78 , 80, 81 View Figures 79–87 , 96, 97 View Figures 88–103

Bryocoris Fallén, 1829: 151   . Type species: Capsus pteridis Fallén, 1807   (by monotypy).


Recognized by the following combination of characters: antennal segment I slightly shorter than head width and distinctly longer than vertex width; labium slightly surpassing fore coxa, labial segment IV short, about twice as long as wide, slightly tapering at apex ( Fig. 23 View Figures 22–33 ); pronotal collar matt, narrow, equal to width of antennal segment I, finely scalloped posteriorly ( Fig. 20 View Figures 10–21 ); pronotum with shallow punctures; sides of scutellum finely scalloped or almost smooth ( Fig. 29 View Figures 22–33 ); costal margin of hemelytron pale yellow, sometimes darkened in apical one-quarter; genital capsule with large twin-coned process above left paramere ( Figs 57 View Figures 46–58 , 62 View Figures 59–78 ); left paramere more than three times as large as right paramere, with distinct sensory lobe and strongly bent apex ( Fig. 69 View Figures 59–78 ); both sexes, sometimes only females, occasionally or usually bracypterous ( Fig. 5 View Figures 1–9 ).

The genus is most similar in body proportions and coloration to Cobalorrhynchus Reuter, 1906   ; however, species of Cobalorrhynchus   clearly differ from species of Bryocoris   in having claw-shaped, long, thin, and strongly tapering labial segment IV, brown line on the costal margin of hemelytron, and by the structure of genital capsule and left paramere.

Diversity and distribution

Bryocoris   includes ten species: Bryocoris bui Hu and Zheng, 2000   ; Bryocoris concavus Hu and Zheng, 2000   ; Bryocoris formosensis Lin, 2003   ; Bryocoris gracilis Linnavuori, 1962   ; Bryocoris insuetus Hu and Zheng, 2000   ; Bryocoris montanus Kerzhner, 1972   ; Bryocoris nitidus Hu and Zheng, 2004   ; Bryocoris persimilis Kerzhner, 1988   ; Bryocoris pteridis (Fallén, 1807)   , and Bryocoris xiongi Hu and Zheng, 2000   . Most of these species are restricted to southern China (Sichuan and Yunnan provinces, Taiwan). Bryocoris persimilis   and B. montanus   are known from the Russian Far East, with B. montanus   also recorded from Japan ( Yasunaga, 2000). The distributional range of B. gracilis   extends from Sichuan and Hunan provinces of China, Taiwan, and Japan to New Guinea. Only B. pteridis   has a wide trans-Palaearctic range, which is almost entirely confined to forest regions.


Relatively little attention had been given to this genus until Hu & Zheng’s (2000) work on the Bryocoris   of China. These authors described four new species, restored Cobalorrhynchus   as a subgenus of Bryocoris   , and provided an illustrated key to species and detailed diagnoses for both subgenera.













Konstantinov, Fedor V. & Knyshov, Alexander A. 2015

Bryocoris Fallén, 1829: 151

Fallen CF 1829: 151