Homocoprini, Génier & Darling, 2024

Génier, François & Darling, James D. G., 2024, Revision of the South American genera Andinocopris new genus and Homocopris Burmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae: Homocoprini new tribe), European Journal of Taxonomy 925, pp. 220-267 : 261-262

Homocoprini tribe nov.

urn:lsid:zoobank.org:act:A86B4671-86B5-41A2-B468-611826BCD7C9

Type genus

Homocopris Burmeister, 1846 , present designation.

Differential diagnosis

The only group of Scarabaeinae with a lateral pronotal carina combined with a setose elytral underside.

Description

Typical tunneler overall body shape. Size moderate to large; secondary sexual characters present and located on dorsum of head and pronotum; integument rather thin and flexible; antennal club elongate oval, symmetrical; internal surface of first segment of antennal club pubescent; distal edge of epipharynx straight; anterior pronotal membrane covering entire width of head insertion; lateral pronotal declivity with a strong carina connected to the lateral pronotal edge anteriorly, surface between carina and lateral edge flat and upright (slightly concave and overlapping only in A. achamas ); posterior portion of pronotal lateral edge at most slightly sinuate; elytra with eight visible striae, stria eight absent or vestigial in some specimens of A. achamas ( Fig. 62 ), stria 9 absent on basal fifth, distinct from stria 10 throughout; elytral internal surfaces with long semi erected setae; tarsal claws present; meso and metatibiae with four edges in mid cross section, expanding from base to apex and lacking transverse ridges on external surface; hindwings with AA3 connected to Cu and RP1 narrow and sharply defined throughout; apodeme of metasternalmetacoxal membrane present; unguinotarsal axillary lamellae present; medial endophallite absent.

Genera included

Andinocopris gen. nov. and Homocopris Burmeister, 1846 .

Figures

Figs 58–64. 58. Copris victorini Boheman, 1857, pronotum oblique view. 59. Sulcophanaeus carnifex (Linnaeus, 1758), pronotum oblique view. 60. Andinocopris achamas (Harold, 1867) gen. et comb. nov., meso-metasternal suture, ventral view.61. Homocopris torulosus(Eschscholtz, 1822), mesometasternal suture, ventral view. 62. A. achamas, left elytron, oblique view. 63. H. torulosus, elytron underside. 64. Sulcophanaeus carnifex, hindwing.