Andinocopris, Génier & Darling, 2024

Génier, François & Darling, James D. G., 2024, Revision of the South American genera Andinocopris new genus and Homocopris Burmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae: Homocoprini new tribe), European Journal of Taxonomy 925, pp. 220-267 : 223

Genus Andinocopris gen. nov.

urn:lsid:zoobank.org:act:DB47A9F1-784F-4431-A7AA-2044987EF94D

Type species

Pinotus achamas Harold, 1867 , present designation.

Differential diagnosis

Dorsal ocular width greater than one quarter of interocular distance. Lateral pronotal carina and pronotal edge joined anteriorly and posteriorly, forming closed ellipse. Pronotal armament in the form of an overhanging ridge or bifurcating projection in large individuals of both sexes. Elytral stria 8 usually absent, remnant present on apical elytral hump and connected to stria five in some individuals of A. achamas ( Fig. 62 ). Meso-metasternal suture posteriorly arcuate ( Fig. 60 ). ♂ FLP endophallite sigmoidal, abruptly narrowing and inflecting medially ( Figs 31–32 ).

Etymology

Andinocopris is a combination of the New Latin adjective ‘ andinus ’ and the Ancient Greek noun ‘κόπρος’ (‘ kópros ’). The former refers to the Andes, where the genus occurs at high elevations from Colombia to Peru, and the latter refers to its association with dung.

Description

Body length 20–34 mm. Anteroventral clypeal tooth, rather small, transverse and varying from truncated to acute in frontal view. Dorsal ocular width greater than one-quarter interocular distance. Anterior pronotal edge straight ( A. achamas ) or slightly sinuous ( A. buckleyi ) medially. Anterior pronotal membrane very wide laterally, transition between membrane and anterior edge of pronotum ill-defined medially. Anteromedial pronotal surface smooth. Lateral pronotal carina and pronotal edge joined anteriorly and posteriorly, forming closed carinated ellipse. Pronotal armament in large individuals of both sexes forming overhanging ridge or bifurcating projection; ventral surface of overhanging armament uneven. Anterior apex of prosternum broadly rounded. Underside of elytron with long semi-erected setae. Elytral stria 8 usually absent, remanent present on apical elytral hump and connected to stria five in some individuals of A. achamas . Hindwing with AP reduced ( Figs 39–40 ). Meso-metasternal suture posteriorly arcuate ( Fig. 60 ). Surface of median metasternal lobe convex, some individuals with shallow and narrow longitudinal sulcus; surface setigerous anterolaterally, impunctate or finely punctate throughout. Pygidial surface finely punctate. ♂ paramere tapering slightly in lateral view; apex blunt. FLP endophallite sigmoidal, narrowing and inflecting centrally.

Distribution

Colombia, Ecuador, Peru.

Figures

Figs 58–64. 58. Copris victorini Boheman, 1857, pronotum oblique view. 59. Sulcophanaeus carnifex (Linnaeus, 1758), pronotum oblique view. 60. Andinocopris achamas (Harold, 1867) gen. et comb. nov., meso-metasternal suture, ventral view.61. Homocopris torulosus(Eschscholtz, 1822), mesometasternal suture, ventral view. 62. A. achamas, left elytron, oblique view. 63. H. torulosus, elytron underside. 64. Sulcophanaeus carnifex, hindwing.

Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling & Génier sp. nov., holotype, ♂ (MZSP).

Figs 37–41. 37–38. Genital segment, dorsal view. 37. Homocopris grossiorum Darling & Génier sp. nov., holotype, ♂ (CEMT). 38. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 39–41. Hindwing, dorsal view. 39. Andinocopris achamas (Harold, 1867) gen. et comb. nov. 40. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov. 41. Homocopris torulosus.