Thiodia klapperichi ( Razowski, 1967 ) Bidzilya & Blackstein & Budashkin & Šumpich & Tsvetkov, 2022

Bidzilya, Oleksiy, Blackstein, Hans, Budashkin, Yuriy, Šumpich, Jan & Tsvetkov, Eugene, 2022, New synonyms and redescription of two little-known Thiodia species (Lepidoptera, Tortricidae): T. ursulana (Kennel, 1919) comb. nov. and T. klapperichi (Razowski, 1967) comb. nov., Zootaxa 5124 (4), pp. 458-470 : 462

Thiodia klapperichi ( Razowski, 1967) comb. nov.

Figs 6–13

Eriopsela klapperichi Razowski, 1967: 95 .

= Eucosma buratana Blackstein, 2014: 66 , syn. nov.

Type material. Holotype of Eriopsela klapperichi , ♂, labelled: “ J. Klapperich, Pagmangebirge, 3000 m, 28.8.53, O. Afghanistan ” | “ Holotype ” | “G.S.-5663 ♂, Eriopsela klapperichi Raz. , det. J. Razowski ” | Prep. Nr. 4882 ♂ ” ( SMNK). Holotype of Eucosma buratana , ♂, labelled: “ 7.x. 1953 г., ТадЖикистан у подноЖЬя г. Буратау, на свет, Ю. Щеткин” [7.x.1953, Tajikistan, near the foot of Buratau mt. , on light, Yu. Shchetkin ”] ( MfN).

Additional material examined. ♂, [ Turkmenistan] Каrа-Каlа , Ashhabad Province, 28.ix.1961 , on light (Krasilnikova); ♂, same data as for preceding but 29.ix.1961 ; ♀, same data as for preceding but 5.x.1961 ; ♂, Phiryuza env., Ashhabad Province, [no data].ix.[no year], on light (Krasilnikova) ; ♂, Badkhyz , Kyzyldzhar, 3.x.1980 (Pechen’) ; ♂, Tajikistan, near the foot of Buratau mt. , on light, 7.x.1953 (Shchetkin) (all ZIN).

Eriopsela klapperichi Razowski, 1967 was described from the male holotype from Pagman Mountains in Afghanistan. The specimen is in rather poor condition ( Fig. 6 ), however the genitalia are characteristic by having a deep sub-ovate emargination on the ventral margin of the valva ( Fig. 12 ). Eucosma buratana was described from two males collected near the foot of Buratau Mountain in Tadzhikistan. The photographs of wings and the male genitalia in the original description ( Blackstein 2014, figs 2, 3) fully agree with the holotype of E. klapperichi . Our study of additional material (five males and one female from Turkmenistan and one male from the type locality of E. buratana in Tajikistan) showed the presence of small rounded bloating on the base of the male forewing ( Fig. 10 ). This character is an autapomorphy of the genus Thiodia ( Kuznetzov 2001: 440) . Taking into consideration that the genitalia of both sexes also agree in general with Thiodia (see remarks in Discussion), we transfer the species to this genus and we propose the new synonymy: Eucosma buratana Blackstein, 2014 , syn. nov. of Thiodia klapperichi ( Razowski, 1967) comb. nov.

Below we provide a description of the female genitalia of T. klapperichi based on a specimen from Turkmenistan ( Fig. 9 ).

Female genitalia ( Fig. 11 ). Papillae anales elongate, membranous, relatively small, covered by fine bristles. Posterior apophyses longer than anterior apophyses, strongly broadened in posterior 1/2, narrowing in middle, very thin in anterior 1/2. Anterior apophyses well sclerotised, slightly bent at their bases. Antrum cup-shaped. Ductus bursae membranous, cingulum long, distinctly sclerotised; long anterior portion of ductus rolled (shown ventrolaterally on fig. 11, so as bursa copulatrix), entirely enclosing the corpus bursae in its original state. Sternum VII broad and short, anterior margin straight, widely rounded on the sides; vane-like, sclerotised areas located posteriorly on the sides, covered by modified bristles (very small, oval). Corpus bursa spherical, membranous, with two horn-like signa located posteriorly opposite to each other; one of these signa very small and hardly noticeable.

Distribution. Afghanistan, Tajikistan, Turkmenistan (new record).

Figures

FIGURES 6–13. Thiodia klapperichi (Razowski, 1967). 6–10. Adults. 6. Holotype of E. klapperichi. 7. Holotype of E. buratana. 8. Male, Turkmenistan. 9. Female, Turkmenistan. 10. Fragment of forewing with bulging (indicated by arrow). 11. Female genitalia. 12, 13. Male genitalia. 12. Holotype of E. klapperichi. 13. Holotype of E. buratana.