Atroposia Pogue

Brown, John W., 2019, New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae), Zootaxa 4671 (2), pp. 195-222: 198-199

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Atroposia Pogue

new genus

Atroposia Pogue  , new genus

Type species: Conchylis oenotherana Riley, 1881  , by monotypy.

Atroposia Pogue  and Mickevich, 1990: 322 (nomen nudum).

Cochylini  New Genus 1 [ oenotherana  group]: Brown 2005: 207; Metzler and Brown 2014: 278.

Atroposia is proposed to accommodate the single species oenotherana  . Originally described in Conchylis Sodoffsky, 1837  (an unjustified emendation of Cochylis  ), the species was assigned to Phalonia Hübner  , [1825], 1816 by McDunnough (1939) and placed in “incertae sedis” by Powell (1983). Pogue (1986) proposed the genus Atroposia for the species in his unpublished Ph.D. dissertation; Razowski (1997) treated oenotherana  under “Unplaced Cochylini  ”; and Brown (2005) and Metzler & Brown (2014) treated it as “ Cochylini  New Genus”. According to Pogue (1986), Atroposia is most closely related to Spinipogon  among North American genera. However, based on facies and genitalia, Atroposia appears to be related to Cochylis  .

Diagnosis. Atroposia is easily distinguished from other genera by facies ( Figs. 3, 4View FIGURES 1–10), which feature a somewhat two-toned forewing with a yellow-tan basal half and a pinkish red distal half with a variably developed yellowish spot. However, in some specimens both the basal and distal halves of the wing are overscaled with dark gray, reducing the two-toned pattern. Putative autapomorphies of Atroposia include the unusual shape of the valva with a distinctly pointed costa and sacculus; the distal portion of the phallus, with a pair of long, pointed, flattened, lateral sheaths and three platelike internal sclerites; and the broad sterigma composed of four fused spheroid plates. The absence of the male hindwing costal roll in A. oenotherana  distinguishes it from most species of Cochylis  and related genera (e.g., Cochylichroa, Cochylidia  , Thyraylia Walsingham, 1897  ).

Description. Head: Vertex and upper frons rough scaled, lower frons with appressed scales; ocellus present; sensory setae of antenna 1.0–1.2 times flagellomere diameter in male, shorter in female; labial palpus porrect, com- bined length of all segments 1.6–1.8 times diameter of compound eye, scaling of segment II concealing segment III; maxillary palpus 1-segmented. Thorax: Posterior crest present; lateral scale tufts of metanotum flat. Forewing in male slightly broadened distally with costa weakly convex and termen straight, moderately oblique, forewing not expanding distally in female, apex rounded in both; forewing length 3.8–5.4 mm, length 2.5–3.4 times width, slightly wider in female; without costal fold in male; Sc less than 0.5 wing length; R 1 originating at or beyond middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating about 0.75 length of discal cell; CuP absent; A1+2 stalked at 0.5 length. Hindwing length 2.5–2.8 times width; costa straight, without costal roll; apex rounded; termen straight, oblique; Sc+R 1 ca. 0.75 wing length, Rs and M 1 stalked ca. 0.65 length of M 1; M 3 and CuA 1 separate; CuA 2 originating 0.65 length of discal cell; male frenulum with one acanthus, female with two. Abdomen: Unmodified. Male genitalia ( Fig. 20View FIGURES 19–26) with tegumen short, broad, convexly rounded at top; uncus absent; socius small, digitate, setose; transtilla a well-developed band, with triangulate to hooklike median process, keeled ventrally; valva highly modified, short, broad basally, with well-developed costal and saccular portions; costal portion broad at base, constricted apically into a robust, curved spine; saccular half with a short, blunt dorsal process and a longer, robust, pointed basal process; inner base of sacculus with elongate, pointed, extension confluent with arms of vinculum; vinculum arms free distally; juxta large, subrectangular, with a pair of dorso-lateral lobes. Phallus large, length about equal to basal width of valva, distally with three median platelike processes covered with minute spicules, subtended laterally by a pair of flattened, distally-pointed flanges extending ventro-terminally; vesica with minute cornuti. Female genitalia ( Fig. 34View FIGURES 33–38) with ovipositor short; papillae anales narrow, weakly sclerotized, setose, joined posteriorly; length of apophyses anteriores ca. 1.2 times length of apophyses posteriores; sterigma large, composed of four fused spheroid plates, sterigma not attached to apophyses anteriores; ductus bursae short, broad, undifferentiated from corpus bursae; corpus bursae a short convoluted sac lacking a signum; accessory bursa originating from ductus bursae.

Etymology. The name comes from the Greek “atropos, meaning without turn. In Greek mythology, Atropos (or Aisa) was one of the three Moirai, the goddess of fate and destiny. The name is feminine in gender.












Atroposia Pogue

Brown, John W. 2019


Metzler, E. H. & Brown, J. W. 2014: 278
Brown, J. W. 2005: 207