Cagiva Pogue

Brown, John W., 2019, New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae), Zootaxa 4671 (2), pp. 195-222: 201

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Cagiva Pogue

new genus

Cagiva Pogue  , new genus

Type species: Phalonia cephalanthana Heinrich, 1921  , by monotypy.

Cagiva Pogue  and Mickevich, 1990: 322 (nomen nudum).

Cochylini  New Genus 2 [ cephalanthana  group]: Brown 2005: 208; Metzler and Brown 2014: 278.

Cagiva is proposed for Phalonia cephalanthana  , a species that has long defied confident generic assignment. The species was described in Phalonia  by Heinrich (1921) and placed in “incertae sedis” by Powell (1983). The monotypic genus Cagiva was proposed by Pogue (1986) in his unpublished Ph.D. dissertation. Brown (2005) and Metzler and Brown (2014) followed that concept, designating the manuscript name Cagiva as “ Cochylini  New Genus”.

Diagnosis. Putative autapomorphies for Cagiva include the prominent, well-sclerotized socii, and the distally bifurcate phallus. The genus is further characterized by a unique combination of characters, some of which are shared with other genera: the 3-segmented maxillary palpi (shared with Thyraylia  , Nycthia  , and few others), the absence of cornuti in the vesica (shared with Eupinivora Brown, 2013  , Honca  , and Nycthia  ), and the connate condition of hindwing veins M 3 and CuA 1 (shared with a few other genera). As in most Cochylina, the male of Cagiva lacks a forewing costal fold and has a well-developed hindwing costal roll.

Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna 0.8–1.0 times flagellomere diameter in male, much shorter in female; labial palpus porrect, combined length of all segments 1.1–1.2 times diameter of compound eye, segment III exposed; maxillary palpus 3-segmented. Thorax: Posterior crest present; lateral scale tufts of metanotum flat. Forewing ( Fig. 2View FIGURES 1–10) length 4.0– 4.7 mm, length 2.4–2.8 times width, slightly wider in female; costa gently arched from base to apex, without costal fold in male, apex abruptly rounded; termen straight; Sc slightly less than 0.5 wing length; R 1 originating beyond middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating ca. 0.65 length of discal cell; CuP absent; A1+2 stalked at ca. 0.5 length. Hindwing moderately broad, length 2.3–2.8 times width; costa straight, with costal roll extending about 0.65 length of costa in male; apex rounded; termen slightly concave below apex; Sc+R 1 ca. 0.25 wing length in male, 0.65–0.75 wing length in female; Rs and M 1 stalked ca. 0.7 length of M 1; M 3 and CuA 1 connate; CuA 2 originating ca. 0.65 length of discal cell; frenulum with one acanthus in male, two in female.Abdomen: Unmodified. Male genitalia ( Fig. 21View FIGURES 19–26) with uncus absent; socius prominent, well sclerotized, setose, distally elongate and curved mesially; transtilla well developed, median process a small rounded tooth; valva length 1.4‒1.5 times width; costa sclerotized, relatively straight; sacculus well defined, confined to basal 0.33 of venter of valva, terminating in a rounded lobe; vinculum arms weakly join distally by thin membrane. Phallus large, ca. 1.3 times length of valva, straight, parallel-sided in basal 0.5, broadened in distal 0.5, bifurcate apically, asymmetric; vesica without cornuti. Female genitalia ( Fig. 35View FIGURES 33–38) with telescopic ovipositor; papillae anales narrow, weakly sclerotized, setose, joined posteriorly; length of apophyses anteriores ca. 2.0 times that of apophyses posteriores, ventral branch of apophyses anteriores reduced, apophyses with rounded swelling at anterior tip; sterigma complicated, comprised of weakly sclerotized sacs, some bearing setae, and a crescent-shaped plate posterad of ostium; sterigma not attached to apophyses anteriores; ductus bursae relatively broad, rather uniform in width, ca. 0.65 as long as corpus bursae, differentiated from corpus bursae, with caudal 0.7 weakly sclerotized; accessory bursa originating near ostium; corpus bursae oblong, covered with minute spicules, signum lacking.

Etymology. The name is an arbitrary combination of letters and is considered feminine in gender.












Cagiva Pogue

Brown, John W. 2019


Metzler, E. H. & Brown, J. W. 2014: 278
Brown, J. W. 2005: 208