Pupinidius pupinidius ( Moellendorff, 1901 )

Wu, Min & Zheng, Wei, 2009, A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species, Zootaxa 2053, pp. 1-31 : 23-25

15. Pupinidius pupinidius ( Moellendorff, 1901)

Figs 1 , 7A–E

Buliminus (Pupinidius) pupinidius Moellendorff, 1901: 345 , pl. 11, figs 6–8; Kobelt, 1902: 847, pl. 121, figs 1–5. Pupinidius pupinidius Yen, 1939: 84 , pl. 7, fig. 51; Schileyko, 1998: 190, fig. 234.

Material examined. ZI RAS-N613: type material, det. Moellendorff, 1885-?, coll. Potanin?. SMF-41992: Lectotype. Dshie-dshou (an unrecogized locality name. Most probably near Zhouqu), Kansu, China. Slg. O.v.Moellendorff. SMF-41993: Paratypes. 3 fmss. Dshie-dshou, Kansu, China.

Distribution. Gansu (type locality).

Type locality. Dshiedshou (Gansu).

Diagnosis. Shell banded beneath suture except in body whorl; faintly striate or not. Aperture with angular tubercle. Most swollen part at penultimate and body whorls. With 6.875–7.375 whorls; height 21.1–23.8, diam. maj. 12.9–13.9. Penis without penial caecum; internally with more than 2 fine longitudinal pilasters. Papilla developed by fused proximal pilasters forming V-shaped structure absent. Epiphallus straight. Epiphallic caecum present.

Re-description of shell. Cylindrical-conic; apex protruded. Most swollen part at penultimate and body whorls. Shell dextral; thin; solid; opaque; glossy; not speckled; not spirally grooved except very weakly grooved at umbilicus region; with 6.875–7.129–7.375 whorls. Whorls not shouldered; slightly convex. Growth lines usually not very clear. Embryonic shell smooth; polished; with 1.500–1.563–1.625 whorls. Postnuclear whorls smooth. Suture normal, without narrow defined zone on beneath whorl. Last whorl gradually ascending towards aperture; rounded at periphery; without smoothed spiral peripheral depression nor rugate region. Shell 21.1–22.2–23.8 high; 12.9–13.4– 13.9 in diam. maj. Height/width ratio 1.58–1.66–1.72. Aperture in a plane; completely attached to body whorl; truncate-ovate; not adnate; not armed; with angular tubercle; 9.9–10.7–11.2 high; 8.5–8.8–9.0 wide. Ratio of shell height to aperture height 1.97–2.06–2.13. Peristome reflexed with distinct cuff. Cuff made by the reflexed peristome more or less curved backward. Peristome expanded; sharp. Parietal callus indistinct. Columellar margin reflexed. Columella vertical. Outer edge of columellar lip vertical. Umbilicus a narrow slit. Apical whorls normally coloured. Shell multicoloured; apex and umbilical region light brownish, with a grey-white spiral band occupying about 2/3 of each whorl just above suture, starting from about fourth or fifth whorl. Aperture white.

Genitalia. See Schileyko (1998: 190, fig. 234).

Figures

FIGURE 1. Distribution map of Pupinidius spp. Red dot, Pupinidius anocamptus, P. nanpingensis nanpingensis and P. nanpingensis ambigua; red circle, P. porrectus; black dot, P. pupinella altispirus; black circle, P. chrysalis; blue dot, P. obrutschewi contractus and P. pupinella pupinella; blue circle: P. latilabrum; green dot, P. melinostoma, P. nanpingensis nanpingensis and P. wenxian sp. nov.; yellow dot, P. obrutschewi obrutschewi and P. streptaxis; yellow circle, P. porrecta pygmaea. P. pupinidius and P. gregorii have not been mapped here due to the unrecognizable locality names. Although the original description of the type locality of P. chrysalis is not accurate and geographically wrong, mapping of the species is possible here based only on the recognized information therein.

FIGURE 7. A – E, Pupinidius pupinidius (Moellendorff, 1901). A, Lectotype, SMF- 41992; B, 1 paratype, SMF- 41993; C – E, type specimens, ZI RAS-N 613. F – G, Pupinidius streptaxis (Moellendorff, 1901). F, type specimen, ZI RAS; G, Lectotype, SMF- 41986. H, Pupinidius wenxian sp. nov., Holotype, HBUMM- 05432 - specimen 1.