Sepsidoscinis Hendel, 1914

Mlynarek, Julia J. & Wheeler, Terry A., 2018, Phylogeny and revised classification of the tribe Elachipterini (Diptera: Chloropidae), Zootaxa 4471 (1), pp. 1-36 : 24-25

Sepsidoscinis Hendel, 1914

( Figs 50–55 ; 57 )

Sepsidoscinis Hendel, 1914: 247 . Type species: Sepsidoscinis maculipennis Hendel, 1914 (original designation).

Diagnosis. Oscinellinae with a dorsoventrally compressed head, an elongated postpronotum, contracted abdomen at base, triangular scutellum with long tubercles, short, patterned wings and long slender arista.

Description. Chloropidae , Oscinellinae . Vertex rounded in lateral view, head dorsoventrally compressed ( Fig. 57 ); frontal triangle shining and bare; frons shining; cephalic chaetotaxy reduced: long peristomal bristles, cruciate postocellar bristles, cruciate short ocellar bristles, long outer vertical bristles, interfrontal setulae long and on margin of frontal triangle, orbital bristles reclinate, of equal size; eye sparsely and microscopically pubescent; gena narrow, vibrissal angle rounded, indistinct, vibrissa with 2 similar bristles; face flat, wide, carina absent; first flagellomere reniform, as long as wide, arista longer than width of frons, slender ( Fig. 50 ); proboscis and palpus short.

Scutum wider posteriorly, as wide as long; thoracic chaetotaxy: 1 anterior, 1 posterior weak notopleurals, 1 postsutural supra-alar bristle, 1 dorsocentral bristle; scutellum dorsally flat, inclined at a 45° angle with scutum, giving impression of a triangular point, usually wider than long ( Figs 52–53 ; 57 ); 1 pair apical scutellar bristles, 1– 2 pairs of lateral scutellar bristles on long tubercles ( Figs 52–53 ); thoracic pleurites bare except for a row of setulae on katepisternum. Wing short, narrow, with a wide dark spot at center ( Fig. 51 ), second costal sector equal to or shorter than third costal sector, distance between crossvein r-m and dm-cu 2.4 times length of dm-cu, anal angle reduced, alula small, much longer than wide; veins pale; halter white. Legs long and slender; male femoral organ absent; small, apical ventral spur on mid tibia; hind tibial spur absent; tibial organ narrow, 0.2 to 0.25 times length of hind tibia.

Abdominal tergites shining, tergites 1+2 very narrow dorsally, tergite 3 enlarged ( Fig. 57 ); sternites slender, with sparse setae; male spiracles 3–5 in membrane near lateral margin of tergite.

Male postabdomen: pregenital sclerites narrow; spiracles 6 and 7 in membrane ventral to lateral margin of dorsal sclerite; epandrium rounded, usually higher than long in lateral view ( Fig. 55 ) and wider than high in posterior view ( Fig. 54 ); surstylus bilobed, upper lobe clavate, lower lobe ending in pointed tip; hypandrium open posteriorly; pregonite fused with postgonite, weakly sclerotized; basiphallus short, weakly sclerotized; distiphallus short, membranous; phallapodeme simple; phallic guide sclerotized; cercus fused with two projections, thin, projecting posteroventrally ( Fig. 54 ).

Female terminalia not modified, cerci separate, cylindrical, setulose.

Geographic distribution. This monotypic genus is restricted to the Oriental realm in southern China, Nepal, India, Sri Lanka, Indonesia and Vietnam.

Remarks. Although Sepsidoscinis is monotypic and is closely related to Anatrichus we have retained it as a separate genus for ease of recognition and nomenclatural stability.

Figures

FIGURES 50–55. Sepsidoscinis maculipennis. 50. Antenna; 51. Wing; 52. Scutellum, dorsolateral; 53. Scutellum, dorsal; 54. Male genitalia, posterior; 55. Male genitalia, lateral. Abbreviations: cer—cercus; epd—epandrium; sur—surstylus. Scale bars—Fig. 51 = 0.5 mm; Figs 54–55 = 0.1 mm.

FIGURES 56–57. 56. Alombus seminitidus: habitus; 57. Sepsidoscinis maculipennis: habitus. Scale bars = 0.5 mm.