Gonaphodiellus sexguttatus ( Schmidt, 1916 )

Dellacasa, Marco, Dellacasa, Giovanni & Gordon, Robert D., 2012, Systematic revision of Gonaphodiellus taxa, with description of two new genera and fourteen new species (Coleoptera: Scarabaeidae: Aphodiinae), Insecta Mundi 2012 (230), pp. 1-41 : 14-15

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Gonaphodiellus sexguttatus ( Schmidt, 1916 )


Gonaphodiellus sexguttatus ( Schmidt, 1916)

( Fig. 62-66 View Figure 57-66 )

Aphodius (Gonaphodiellus) sexguttatus Schmidt, 1916: 114 ; Dellacasa 1988: 240.

Aphodius sexguttatus ; Blackwelder 1944: 213.

Gonaphodiellus sexguttatus ; Skelley et al. 2007: 4.

Type locality. Pichinde, Colombia.

Type repository. Naturhistoriska Riksmuseet, Stockholm, Sweden (type examined).

Redescription. Length 4.5-5.0 mm; elongate, moderately convex, subshiny, glabrous. Dark brownish; clypeal margin and fore angles of pronotum reddish; elytra yellowish spotted basally on second and third interstria and at shoulder, posteriorly with more or less elongate and widened blackish margined yellowish spots on second, fourth, sixth and seventh interstria; legs brownish-red; antennal club dark brown. Head with epistome slightly gibbous on disc, superficially alutaceous, rather finely, evenly, not closely punctured; punctation medially finer and more superficial; clypeus subtruncate anteriorly, round at sides, rather finely bordered, edge glabrous; genae round, almost imperceptibly bristled, not protruding from the eyes; frontal suture finely impressed, not tuberculate; front evenly, not closely punctured. Pronotum transverse, moderately convex, superficially alutaceous, somewhat irregularly, not very densely punctured; punctation denser and coarser on sides, sparser and more superficial on disc; lateral margins feebly arcuate, rather finely bordered, edge glabrous; hind angles obliquely truncate, truncation distinctly sinuate inwardly; base bisinuate. Scutellum flat, finely alutaceous, finely sparsely punctured. Elytra convex, almost subparallel-sided, finely striate; striae superficially punctured, faintly crenulate; interstriae feebly convex, very finely sparsely punctured, glabrous, preapically finely alutaceous thus rather dull. Hind tibiae upper spur shorter than first tarsal segment; first tarsal segment almost as long as following four segments combined. Male: pronotum not narrowed anteriorly and relatively somewhat less densely and less coarsely punctured; metasternal plate medially moderately excavate; aedeagus Fig. 65-66 View Figure 57-66 . Female: pronotum somewhat narrowed anteriorly and relatively more densely and more coarsely punctured; metasternal plate almost flat.

Remarks. The species is redescribed on Costa Rican specimens, the characters of which match almost perfectly those of the sole female type from Colombia.

Material examined. COLOMBIA: Pichinde (female holotype, NHRS) . COSTA RICA: Alajuela Prov., Penas Blancas River Valley , m 500-1000, 06.V.1985, leg. Lyon B. (1 ex., CNCI) ; Atlantic Slope , 6 Km N Santa Elena Punt., 31.V.1979, leg. Howden H. & A. (1 ex., CNCI) ; Cartago Prov., Pan Am Hwy, Km 45, 6 Km NW El Empalme, 08-26.VI.1997, leg. Peck S. & J., FIT (3 exx., CNCI) ; idem, m 1975, 08- 26.VI.1997, FIT, leg. Peck S. & J. (1 ex., DCGI) ; La Cañada T. Bi , m 2300, 15.IX.1940, leg. Bierig A. (1 ex., USNM) ; Cartago Prov., Navarro , 16-31.VII.1962 (4 exx., USNM) ; Monteverde , m 1520, 11- 18.VI.1983, leg. Howden H. & A. Howden, FIT (4 exx., CNCI) ; idem, 23-30.VII.1983, leg. Lindeman D. H., FIT (1 ex., DCGI) ; Puntarenas, Monteverde , 26.V-03.VI.1984, leg. Riley E., Rider D. & Ledoux D. (1 ex., DCGI) ; idem, m 1400, 04.VI.1979, leg. Howden H. & A. (1 ex., CNCI) ; idem, m 1500, 23- 27.XI.1991, leg. Howden H. & A., dung traps (1ex., DCGI) ; Punt [arenas]., Monteverde Reserve , m 1500, 22.VIII.1967, leg. Howden H. & A. (8 exx., CNCI) ; San José, 05.VIII.1910, leg. Schmidt H. (1 ex., NMPC) ; S[an]. J[osé]., 9 Rd Km S of Division, 19.V.1991, leg. Curoe D. (2 exx., USNM) ; 8 rd km N Vara Blanca, Volcan Poas, 1500 m, Alajuela Prov., 11.V. 1985, leg J. Doyen, Black light (2 exx., EMEB) . EL SALVADOR: Monte Cristo , 23 Km N Metapán, m 2300, 08-10.V.1971, leg. Howden H. F. (2 exx. CNCI, DCGI) ; Monte Cristo , 07-09.V.1958, leg. Cartwright O. L. (1 ex., USNM) ; Hda. Montecristo, Miramundo, Metapán , m 2300, 08.V.1971, leg. Steinhauser S. & L., cloud forest (1 ex., FSCA) . HONDURAS: F. Morazán, 30 Km E Tegucigalpa, Cerro Hyuca , m 1800, 10.VI.1994, leg. Howden H. & A. (122 exx., CNCI, DCGI) ; Paraíso, Cerro Monserrat , 7 Km SW Yuscarán, m 1800, 24.IV.1994, leg. Howden H. & A. (10 exx., CNCI, DCGI) ; Depto. Paraíso, Monserrat Mt. , 01.IV.1994, leg. Ganaway R. (1 ex., DCGI) ; El Paraíso, Cerro Monserrat , 6.9 mi. W Yuscaran, m 1760, 07.VI.1994, leg. Anderson R. S., cloud forest litter, Berlese (2 exx., CNCI) ; Ocotepeque, R. B. Gulsayote, 22.V.1995, leg. Turnbow P., mv+bl (10 exx., FSCA; 2 exx., DCGI) ; Dept. Olancho, La Muralla Pk. N., 24-27.V.1995 ; leg. Huether J. (1 ex., FSCA) Olancho, Parque Nacional La Muralla , 15°05’49"N- 86°44’17"W, m 1450, 04-07.VII.2002, leg. Smith A. T. B. & Ocampo F. C. (34 exx., FSCA; 2 exx., DCGI) GoogleMaps . MEXICO: Chiapas: Mpio. Angel Albino Corzo, Reserva El Triúnfo, rd. to Finca Prussia , nr. campamento, 15°39’44.7"N- 092°48’40.7"W, 25.IX-25.X.2002, leg. Galvez R. S., FTT (1 ex., FSCA) GoogleMaps . PANAMA: Chiriqui, Hartmann’s Finca, Sta. Clara , 15-18.VI.1985, leg. Riley & Rider (1 ex., DCGI) .

Distribution. Colombia, Costa Rica, El Salvador, Honduras, Mexico (Chiapas), Panama.

Bionomics. Poorly known; the majority of specimens examined were collected in spring.


Swedish Museum of Natural History, Entomology Collections


Canadian National Collection Insects


Smithsonian Institution, National Museum of Natural History


National Museum Prague


Florida State Collection of Arthropods, The Museum of Entomology














Gonaphodiellus sexguttatus ( Schmidt, 1916 )

Dellacasa, Marco, Dellacasa, Giovanni & Gordon, Robert D. 2012

Gonaphodiellus sexguttatus

Skelley, P. E. & Dellacasa M. & Dellacasa G. & R. D. Gordon 2007: 4

Aphodius sexguttatus

Blackwelder, R. E. 1944: 213

Aphodius (Gonaphodiellus) sexguttatus

Dellacasa, M. 1988: 240
Schmidt, A. 1916: 114