Arpactophilus arhoe, Breitkreuz, Laura C. V., Ohl, Michael & Engel, Michael S., 2016

Breitkreuz, Laura C. V., Ohl, Michael & Engel, Michael S., 2016, A review of the New Caledonian Arpactophilus (Hymenoptera: Crabronidae), Zootaxa 4063 (1), pp. 1-66 : 14

publication ID

https://doi.org/ 10.11646/zootaxa.4063.1.1

publication LSID

lsid:zoobank.org:pub:80401ED8-C6BA-4420-9109-854C5CC1E88D

DOI

https://doi.org/10.5281/zenodo.6088293

persistent identifier

https://treatment.plazi.org/id/03E4FF4A-FF94-FFCE-3FB9-FF00C33EF9FF

treatment provided by

Plazi

scientific name

Arpactophilus arhoe
status

sp. nov.

Arpactophilus arhoe View in CoL sp. nov.

( Figs. 29–30 View FIGURES 29 – 34 )

Diagnosis. Arpactophilus arhoe is the only New Caledonian species in the genus with the combination of a black clypeus ( Fig. 29 View FIGURES 29 – 34 ), a continuous occipital carina ( Fig. 5 View FIGURES 5 – 9 ), an imbricate and punctured gena without a carina, a black metasoma, and a bulge on sternum II ( Fig. 30 View FIGURES 29 – 34 and as in Fig. 2 View FIGURES 1 – 4 ).

Description. FEMALE: Total length 5.4 mm; forewing length 3.4 mm.

Body black, with areas of dark yellow and brown. Dark yellow: labrum; palpi; scape; pedicel; flagellum in front; anterior pronotal margin; tegula; coxae apically; trochanters; most of femora; tibiae; tarsi. Brown: mandible partially; posterior surface of of flagellum; pronotal lobe apically; most of coxae; femora partially ( Fig. 30 View FIGURES 29 – 34 ). Wings hyaline; pterostigma dark brown.

Head about 0.9 × as long as wide in frontal view. Apical margin of clypeus medially projecting, with shallow, broad medial incision ( Fig. 29 View FIGURES 29 – 34 .). Apical margin of labrum with two small lobes. Ventral mandibular tooth about ¼– 1/5 of total mandibular length, not reaching opposite mandibular base. Palpal formula 5:4. Frons imbricate, with dense, coarse punctation, punctures separated by less than a puncture width and dense setae ( Fig. 29 View FIGURES 29 – 34 ). Frontal carina present from median ocellus to almost apical margin of clypeus, elevated to a distinct ridge on clypeus. Scape 3.9 × as long as wide. Ocellar triangle anterior of tangent between upper posterior orbits of compound eyes, lateral ocelli anterior of tangent by less than their diameter. Occipital carina continuous ( Fig. 5 View FIGURES 5 – 9 ). Gena imbricate, with dense, coarse punctation and dense setae, without tubercles, spines, or dorsoventral carina. Hypostomal midventral line carinate, with bordering short transverse carinulae on hypostomal integument, not angulate.

Mesosoma about 1.7 × as long as wide in dorsal view. Propodeum about 0.8 × as long as wide in dorsal view. Mesosoma imbricate with additional dense punctation ( Fig. 30 View FIGURES 29 – 34 ), except dorsal surface of propodeum; lateral surface of propodeum also transversely carinulate; dorsal surface of propodeum coarsely reticulate (similar to pattern in Fig. 18 View FIGURES 16 – 22 ). Pitted sulcus present posterior of fine mesoscuto-scutellar sulcus. Hypersternal sulcus long and pitted. Metafemur 2.6 × as long as wide. Metatibia without differently colored area apically. Pretarsal claws without teeth. Forewing with two submarginal cells; anterior border of submarginal cell II slightly shorter than posterior border. Hind wing with two distal hamuli.

Metasoma polished, with sparse punctation ( Fig. 30 View FIGURES 29 – 34 ). Metasomal sternum II swollen medially, forming bulge (as in Fig. 2 View FIGURES 1 – 4 ). Pygidium with broad row of silk setae apically.

MALE: Unknown

Remarks. Arpactophilus arhoe can be distinguished from all other Arpactophilus with a bulge on the metasomal sternum II by the form of its clypeus. In A. arhoe , the apical margin of the clypeus is medially projecting, with a shallow, broad medial concave incision. Among species with a bulging metasomal sternum II, the form of the clypeus is closest to that of A. arha , but the two species differ noticeably in coloration, especially of the clypeus and lower frons, which are black in A. arhoe ( Fig. 29 View FIGURES 29 – 34 ) and yellow in A. arha ( Fig. 27 View FIGURES 23 – 28 ). Arpactophilus pwaamei and A. tayo are also similar in coloration and clypeal form, but they have no bulge on metasomal sternum II.

Material examined. HOLOTYPE ♀: “Nouvelle Calédonie, Pinda 30m, 18.IV.1995, Réc. Chazeau & Jourdan [ MNHN].

Etymology. The specific epithet is taken from the name Arhö , one of the New Caledonian native languages. It is treated as a noun in apposition.

MNHN

Museum National d'Histoire Naturelle

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