Metaconchoecia, Chavtur, Vladimir Grygorjevich, 2003
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publication ID |
https://doi.org/ 10.5281/zenodo.156688 |
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DOI |
https://doi.org/10.5281/zenodo.6276951 |
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persistent identifier |
https://treatment.plazi.org/id/03E51C2A-FFC0-FFF0-FEE0-FB0B8398FAEC |
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treatment provided by |
Plazi |
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scientific name |
Metaconchoecia |
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Key to species of “ skogsbergi ” complex
Only for male
1. Copulatory appendage is very curved .......................................................... M. australis View in CoL
Copulatory appendage is straight or almost straight ..................................................... 2
2. Left asymmetrical gland is placed more than17% length of carapace behind tip of rostrum .................................................................................................................... M. discovery
Left asymmetrical gland is placed less or about 15% of carapace behind tip of rostrum ....................................................................................................................................... 3
3. Length is about or more than 1.7 mm, dorsal margin is broadly Vshaped with apex just posterior to insertation of protopodite of 2nd antenna, length of setag is about 2.5 length of 1st endopodite segment of 2nd antenna ................................ M. ampla View in CoL nov. sp.
Length is less 1.7 mm, dorsal margin is straight, length of setag is 3 times as long as length of 1st endopodite segment of 2nd antenna........................................................... 4
4. Length is more than 1.3 mm .......................................................................................... 5
Length is less 1.3 mm (except in M.fowleri Form A View in CoL ) ................................................... 7
5. Right asymmetrical gland opens near posterior dorsal corner ..................................... 6.
Right asymmetrical gland is placed somewhat below posterior dorsal corner (about 10% length of posterior margin) ................................................. M. skogsbergi nov.sp.
6. Carapace is short and round in lateral view (height more than 50% of length), setac of 1st antenna is thick, shorter than total combined length of segments 3 to 5, setae of 2nd antenna is large, stout and clawlike ...................................................... M. alta View in CoL nov. sp.
Carapace is more cylindrical (height usually less than 50% of length), setac of 1st antenna is thin and subequal to total combined length of segments 3 to 5, setae of 2nd antenna is tiny, thin, and peglike .................................................. M.longiseta View in CoL nov. sp.
7. Setac of 1st antenna is subequal to total combined length of segments 3 to 5, setae is twice as long as this limb ................................................................................ Mwolferi
Setac of 1st antenna is shorter than total combined length of segments 3 to 5, setae is less than length of this limb .......................................................................................... 8
8. Setaa of 1st antenna is proximally noticeable swollen and extends beyond proximal end of limb ................................................................................... M. crassiseta View in CoL nov. sp.
Setaa of 1st antenna is either usual, or proximally slightly swollen and barely reaches or does not reaches of proximal limb............................................................................ 9
9. Carapace is unusually broad in ventral view (more than 50% of length) ................... 10
Carapace is less broad in ventral view (less than 50% of length), except in M. acuta View in CoL (48.6+ 2.6%)................................................................................................................ 11
10. Capitulum of frontal organ is broad, setac of 1st antenna is short; setae of 2nd antenna is developed, its proximal spines are short .......................................... M. inflata inflata View in CoL
Capitulum of frontal organ is narrow, setac of 1st antenna is short; setae of 2nd antenna is minute, its proximal spines are long ...................................... M. inflata lata View in CoL nov. spp.
11. Setae of 1st antenna is with short spines ......................................................... M. obtusa View in CoL
Setae of 1st antenna is with long spines...................................................................... 12
12 Length is more than 0.9 mm ........................................................................................ 13
Length is less than 0.9 mm ......................................................................... M.rotundata View in CoL
13. Setac of 1st antenna is about half of total combined length of segments 3 to 5, and setaa reaches proximal end of limb............................................................................ 14
Setac of 1st antenna is barely shorter than total combined length of segments 3 to 5, and setaa does not reach proximal end of limb ......................................... M. subinflata View in CoL
14. Length is 1.1 mm or over, setae of 2nd antenna is armed with 1215 pairs of spines..... ....................................................................................................................... M. fowleri View in CoL
Length is less than 1,0 mm, setae of 2nd antenna is armed with 1012 pairs of spines .. ............................................................................................................................ M.acuta View in CoL
Only for female
1. Left asymmetrical gland is situated more than or about 17% length of carapace behind tip of rostrum............................................................................................ M. discoveryi View in CoL
Left asymmetrical gland is situated less than or about 15% length of carapace behind tip of rostrum ................................................................................................................ 2
2. Exopodite of 2nd antenna is about 80% length of protopodite ...................................... 3
Exopodite of 2nd antenna is less than or about 70% length of protopodite ................... 4
3. Length is less than 1.3 mm, height is more than 50% of length, terminal setae on endopodite of 2nd antenna is half as long as exopodite .............. M. inflata lata View in CoL nov.ssp.
Length is more than 1.6 mm, height is less than 50% of length, terminal seta on endopodite of 2nd antenna is about 80% of exopodite ......................... M. ampla View in CoL nov.sp.
4. Terminal setae on endopodite of 2nd antenna is about 40% and 60% length of proto podite and exopodite ......................................................................................... M.acuta View in CoL
Terminal setae on endopodite of 2nd antenna is more than or about 50% length of pro topodite and more than 60% length of exopodite ......................................................... 5
5. Length is less than 0.9 mm ......................................................................... M. rotundata View in CoL
Length is more than 0.9 mm .......................................................................................... 6
6 Length is about or more than 1.3 mm ........................................................................... 7
Length is less than 1.3 mm (except M. fowleri Form A View in CoL ) .............................................. 9
7. Right asymmetrical gland open near posterior dorsal corner........................................ 8
Right asymmetrical gland open somewhat below posterior dorsal corner (about 10% length of posterior margin) ........................................................................ M.skogsbergi
8. Length is about 1.3 mm, 2nd segment of 1st antenna is without seta. M. aff. skogsbergi
Length is more than 1.4 mm, 2nd segment of 1st antenna is armed with minute seta .. 15
9. Setae on 1st antenna is about or more than 2.5 length of limb, terminal seta of 2nd antenna is about or more than 80% length of exopodite ............................................. 10
Setae on 1st antenna is less or about twice as long as limb, terminal setae of 2nd antenna is less than 80% length of exopodite ............................................................. 11
10 Length is about or 1.1 mm or over ................................................................. M. fowleri View in CoL
Length is about or less than 1.0 mm ................................................................ M. obtusa View in CoL
11. Exopodite of 2nd antenna is about 80% length of protopodite ...... M. iflata lata nov. sp.
Exopodite of 2nd antenna is about or less than 70% length of protopodite ................. 12
12. Setae of 1st antenna is twice as long as limb or sensory setae.................................... 13
Setae of 1st antenna is less than twice length of limb or sensory setae .......................... ...................................................................................................... M. crassiseta View in CoL nov. sp.
13. Height and breadth of carapace are more than 50 and 40% of its length, respectively .. ...................................................................................................................... M. australis View in CoL
Height and breadth of carapace are about or less than 50 and 40% of its length, respec
tively .................................................................................................................... 14 14. Carapace is prolonged, capitulum of frontal organ is straight and with rounded tip ...... ........................................................................................................................ M. wolferi View in CoL
Carapace is high, capitulum of frontal organ is curved and with pointed tip ................ .................................................................................................................... M. subinflata View in CoL
15. Capitulum of frontal organ is fused to shaft or separated by indistinct suture, 1st segment of 1st antenna is bare, length of 1st endopodite segment on maxilla is about 1.5 of its height ................................................................................................. M. alta View in CoL nov. sp.
Capitulum of frontal organ is separate from shaft by distinct suture, 1st segment of 1st antenna is armed with dorsodistal spines, length of 1st endopodite segment on maxilla is subequal to height ....................................................................... M. longiseta View in CoL nov. sp.
Order HALOCYPRIDA Dana, 1853 View in CoL
Suborder HALOCYPRIDINA Dana, 1853
Superfamily HALOCYPRIDOIDEA Dana, 1853
Family HALOCYPRIDIDAE Dana, 1853
Subfamily CONCHOECIINAE Müller, 1906
Genus METACONCHOECIA Kock, 1992
Metaconchoecia longiseta nov. sp.
( Figs. 213 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 View FIGURE 11 View FIGURE 12 View FIGURE 13 )
Conchoecia skogsbergii: Chavtur, Shornikov, 1974: 286 (part).
Metaconchoecia skogsbergii: Chavtur, 1976: 105 106 (part); 1977a: 145146 (part); 1977b: 30 (part); 1977c: 20 (part).
Metaconchoecia View in CoL species nov. 2 Chavtur, 1991: 46 (part); 1992: 49, table 2.
Etymology. The specific name “ longiseta ”, is derived from the Latin word “ longa” meaning long and “seta” refers to the exceptionally long cbristle on the endopodite of the second antenna.
Holotype. IBM 2792 – adult male, length 1.42 mm, appendages mounted on slide and valves in alcohol, in collection of the Museum of Institute Marine Biology, Vladivostok, Russia (together with paratypes).
Typelocality. RV Vityaz 39th Cruise, station 5617, sample 126, at 45º49’N, 153º33’E, depth 500 200m, August 5 1966, using a Bogorov – Rass’s Net (S = 1,0 m²).
Paratypes. IBM 2793 –adult male, IBM 2794 – adult male, IBM 2795 – adult male, IBM 2796 – adult female, IBM 2797 – adult female, IBM 2798 –adult female, IBM 2799 – adult male, same station and sample as for holotype. Paratypes IBM 2793 –2799: append MALE
Carapace ( Figs. 2 View FIGURE 2 AF; 3 A; Table 1 View TABLE 1 ). The length of adult specimens ranges from 1.30 1.55 mm. Carapace is relatively elongated. Its maximum height is 4551% of the carapace length (CL), and occurs posterior to the midline. Hence the posterior half of the shell is somewhat larger than the anterior half. The maximum width of the carapace is 4551% CL and is positioned just anterior to midlength; generally the anterior region of the carapace is slightly broader than the posterior region. The shoulder vault is weakly developed and rounded. The dorsal margin is either straight or broadly Vshaped with the apex just posterior to the insertion of the protopodite of the second antenna. The posterior dorsal corner is a rounded angle with a distinct apex. The posterior margin curves evenly, and the ventral margin is almost straight or slightly curved, curving smoothly into posterior end. The rostrum is minute and has a pointed tip. The rostral incisure is fairly shallow. In ventral view, the sides of the carapace are only gently curved. On the right carapace valve the asymmetric gland opens almost at the posterodorsal corner, whereas the gland on the left valve is located at a position just posterior to the rostral incisure. Dorsomedial glands are weakly developed. The glands along the anterior margin of the shell are rather distinct. There is no surface ornamentation.
AC lateral view of left valve of carapace; D lateral view of right valve of carapace; E anterior of right carapace valve; F posterodorsal region of carapace; G distal end of frontal organ.
A ventral view of carapace; B upper lip; C frontal organ and 1st antenna; D terminal part of 1st antenna; E and F second antenna; G – bseta of endopodite of second antenna.
AC medial part of c, d and esetae of 1st antenna; D and E endopodite of left second antenna; F endopodite of right second antenna; G and H hook appendage of endopodite of left second antenna.
Frontal organ ( Figs. 2 View FIGURE 2 G; 3 C). The shaft extends beyond the end of the second segment of the first antenna. The proximal half of the capitulum is expanded, with a strongly convex ventral margin, whereas the distal part is narrow, and either tapers slightly or is parallelsided. The surface of the capitulum is densely covered with numerous minute spines.
First antenna ( Figs. 3 View FIGURE 3 C, D; 4 AC). The segmentation is fairly distinct. The aseta extends back parallel to the limb (except proximally where it loops down and is slightly swollen) not quite reaching its proximal boundary. The bseta is slightly longer than the dseta and bears 911 closely spaced various sized anterior spines and has 57 short spines and followed 56 long closely spaced spines on its posterior margin. The c seta is relatively thin and long, it length being similar to the total length of the third to fifth segments of the limb. The dseta is armed with 89 closely spaced varioussized anterior spines. The longest eseta is about one and a half (or somewhat longer) the length of the limb. The eseta armature comprises 1315 pairs of spines, which lie at an acute angle to the seta. The first segment of the limb contains some dark pigment spots.
Second antenna ( Figs. 3 View FIGURE 3 E, F; 4 DH; 5 AE; 6 A, B). The protopodite has some dark pigment spots. The exopodite is about 60% (or somewhat longer) the length of the protopodite The first endopodite segment bears three triangular ridges. The distoventral surface of this segment is covered with fine hairs. The processus mamillaris is bulbous with a beaklike extension pointing slightly forwards. The bseta is about twice the length of the aseta and 1.5 the length of the 2nd endopodite segment. The c and dsetae are relatively long. The eseta is tiny. The hook appendage on the right limb is strongly developed with a long curved distal section and a number of subterminal ridges. On the left limb the hook appendage is smaller, with a relatively short straight distal section and no subterminal ridges, but it has terminal verruca. The sensory h, j and isetae are about 40% the length of the gseta. The sensory h, j and isetae are about 30% the length of the gseta. The gseta is 8085% the length of the protopodite, about 1.2 the length of the exopodite and 4 times as long as the first endopodite segment. The f and gsetae are somewhat flattened distally.
AD hook appendage of endopodite of right second antenna; E distoventral region of endopodite of right second antenna; F mandible.
A and B end of hook appendage of endopodite of right second antenna; C medial region of main seta of terminal segment of mandible; D basal endite of mandible; E toothed edge and proximal and distal tooth rows of mandibular coax; FH masticatory pad and toothrows of coxa of mandible.
A and B maxilla, endite and terminal region; C – 5th limb.
Mandible ( Figs. 5 View FIGURE 5 F; 6 CH; 8 A, D). The coxale cutting edge has a straight anterior section followed by 1215 teeth. The distal list has a large double pointed posterior tooth, followed by about 25 small teeth. The proximal list is armed with large pointed posterior tooth, 34 small teeth, a second large tooth followed by about 2025 small teeth, one of which near the middle, is larger than the others; the inner surface of this list is covered with very small teeth or papillae. The masticatory pad bears four narrow rounded flaps (covered with exceedingly fine, rather short and exceedingly dense spines), four stout teeth, numerous filaments (with bifurcated tip) and hairs. The cutting edge of the basale has two tube teeth, the posterior one pointed and with spines, the anterior one rounded and bare; these are followed by six serrated teeth, of which the most posterior lacks or almost lacks secondary cusps. The anterior inner tooth is rounded and lacks serrations. The epipodite is triangular and developed. The exopodite is represented by slight and plumose seta. It is as long as the first endopodite segment. The dorsal seta on this segment is rather thin, plumose and subequal to the segment. The longest seta from the three dorsal setae on the second segment is stout and somewhat longer than the main clawlike seta of this limb.
Maxilla ( Figs. 7 View FIGURE 7 A, B; 8 C, D). The most anterior seta on the endite of the precoxale is relatively short and bears very thick secondary setae distally. Its main clawlike seta is pectinate. There are three posterior tube setae, two of which are armed with long spines and the third is bare. On the posterior process of the coxal endite the most anterior tooth has 2 6 stout spines in the middle. The basale has no seta. The first endopodite segment is very broad. Its length and width are subequal. At about the middle of the anterior side there are three shorthaired setae. The posterior side bears four shorthaired setae. The distomedial seta extends just beyond tip of the limb. On the ventral surface of this segment there are about 10 short spines. The terminal segment is comparatively short and wide. The anterior claw is about 1.5 times the length of the posterior claw.
Fifth limb ( Fig. 7 View FIGURE 7 C). The epipodial plate carries four plumose setae in each of the distal, middle and proximal groups of setae respectively and is also armed with some short spines. The first endite of the protopodite bears one long plumose and one short shorthaired setae, and the second endite carries one long plumose and two short shorthaired setae. On the endopodite there are nine setae, two of which are clawlike, threelong and plumose, and remainder are short with shorthairs. The first exopodite segment bears seven ventral, one lateral and one dorsal shorthaired setae. The second segment has two ventral and one dorsal shorthaired setae. The third segment has two fairly long clawlike setae and a shorter ventral seta.
Sixth limb ( Fig. 9 View FIGURE 9 A). The distal, middle and proximal groups of setae of the epipodial plate consist of five, five and six (and one additional short) long and plumose setae respectively and the plate is also covered with some short spines. The longer seta on the endopodite is plumose. On the first exopodite segment there are three short ventral, one plumose long lateral setae and distal row of hairs. The second segment has a minute ventral seta, and the third segment is with a minute ventral seta and longer dorsal seta lying parallel to the limb. The terminal setae of the limb and the exopodite are subequal.
Seventh limb ( Fig. 8 View FIGURE 8 E). The terminal segment bears two setae, one about four times the length of the other.
A and B anterior of proximal toothrow of mandibular –oxa; C and D endites of maxilla; E – 7th limb; F copulatory appendage.
A sixth limb; B copulatory appendage and caudal furca; C distal end of copulatory appendage.
Caudal furca ( Fig. 9 View FIGURE 9 B). There are eight pairs of clawsetae and an unpaired seta dorsal to the smallest pair of clawsetae. The inner surface is covered with fine hairs.
Copulatory appendage ( Figs. 8 View FIGURE 8 F; 9 B, C; Table 1 View TABLE 1 ). The limb is relatively wide and of average length ( Table 1 View TABLE 1 ). Its end is obliquely truncated. The anterior side is distinct concave near the end. There are six or seven oblique muscle bands.
Upper lip ( Fig. 3 View FIGURE 3 B). The posterior ventral edge is interrupted by a deep Vshaped notch. On each side of the notch there are some short flaccid spinelike processes, which decrease in stoutness medially.
FEMALE
Carapace ( Fig. 10 View FIGURE 10 AF). The lengths of adult females range from 1.40 to 1.60mm (mean 1.49mm). The maximum height of the shell is 4753% of its length, and its the posterior part is considerably larger than the anterior part. Its maximum width (3950% CL) occurs at or just posterior to the midline. In all other respects it is similar to the carapace of the males.
Frontal organ ( Figs. 10 View FIGURE 10 GJ; 11 A; 13 A). The shaft extends well beyond the end of the fist antenna and is about double the combined lengths of the 1st and 2nd segments. The capitulum is separated from the shaft by distinct suture. It is rather expanded proximally and tapers to a pointed tip. Its dorsal surface is either straight or slightly concave near the middle. Its ventral surface is usually slightly concave distally. The end is usually downturned, and the surface of the capitulum is partially covered with minute spines.
First antenna ( Figs. 11 View FIGURE 11 A, B; 13 A). The boundary between second and third segments is indistinct. The first segment bears about 10 dorsodistal spines. There is minute dorsal seta (distally spinous) on the second segment. The a, b, c, and dsetae are about half the length of the eseta and are subequal or slightly shorter in length than the limb. The b, c and dsetae have thin bases but the aseta has a thickened base. The eseta is about 1.7 the length of the limb and bears short anterior and long posterior spines placed along its middle and distal sections. It tapers to a pointed tip, which is unflattened.
Second antenna ( Figs. 11 View FIGURE 11 C, D; 12 A, B; 13 B). The protopodite contains some dark pigment spots. The exopodite is short and about 65% the length of the protopodite. The distoventral corner of the first endpodite segment has three triangular ridges and is covered with fine hairs. The aseta is about half length of the bseta. Neither a c nor a dsetae are present. The f, g, h, i and j setae are subequal, about as long as or somewhat shorter than half the length of the protopodite, about 75% of the exopodite, and twice (or somewhat more) as long as the first endopodite segment. They are parallelsided and have pointed tips. The processus mamillaris on each endopodite is similar to the male’s.
Mandible ( Fig. 12 View FIGURE 12 C). It is same as in the male, only somewhat smaller.
Maxilla. Similar to the male’s.
Fifth limb ( Fig. 11 View FIGURE 11 E). Similar to that of the male except there is a plumose lateral seta on the first exopodite segment. All setae are somewhat stouter than those of males.
Sixth limb ( Fig. 13 View FIGURE 13 C). The limb is same as in the male, but differs in following respects: the first exopodite segment has five ventral (three plumose) setae and one dorsal, plumose seta. The setae on the second and third segments are relatively long. The main terminal seta is somewhat shorter than the epipodial plate of this limb.
Seventh limb, Caudal furca ( Fig. 13 View FIGURE 13 D) and Upper lip are all similar to those of males.
A frontal organ and 1st antenna; B medial section of eseta of 1st antenna; C and D anteroventral region of 1st endopodite segment of second antenna; E – 5th limb.
A second antenna; B endopodite of second antenna; C tooth rows and masticatory pad of coxa of mandible.
A medial region of frontal organ and second segment of first antenna; B anteroventral region of endopodite of second antenna; C – 6th limb; D caudal furca.
Remarks. This species is closely related to M. skogsbergi ( Iles, 1953) . Some distinguishing characteristics between the two species are listed in the Table 2 View TABLE 2 .
. .....continued on the next page Characteristic M. skogsbergi M. longiseta nov. sp. M. alta nov. sp.
Female. Frontal organ:
capitulum fused to shaft separated by distinct fused to shaft or sepasuture rated by undistinct suture
First antenna:
dorsal seta absent present present
1st segment bare with dorsodistal spines bare Distribution. This is interzonal species since it occurs both in the epi and mesopelagic zones. M. longiseta View in CoL is widely distributed in the temperate and adjacent subtropical waters between 37º and 61ºN at the depth range 251500m in the NorthWestern Pacific and between 40º and 60ºN from 50 to depth 2500m in the NorthEastern Pacific ( Fig. 1 View FIGURE 1 ). While it has been found in the Sea of Okhotsk and the Bering Sea, it appears to be absent from the Sea of Japan.
Its vertical distribution in the NorthWestern Pacific has been studied in the region of the KurileKamtchatka Trench (Table 3), where M. longiseta View in CoL occurs predominantly in the depth range 300 to 1000 m (single specimens were caught at 100200 and 10001500 m). It is most abundant and its biomass was higher at depths of 300500 m; its abundance declined abruptly below 750 m. Its abundance as a percentage the total halocyprid population is quite small. Its maximum relative abundance of about 4% was in 300500 and 500 750m depth layers, and its highest relative biomass, only ~2%, was at 500 750m. Males, females and juveniles were most numerous in the 300500m layer ( Table 5 View TABLE 5 A).
TABLE 3. Abundance and biomass1 of halocypridids in area of the KurileKamtchatka Trench (RV Vityaz 39th Cruise, 1966; BogorovRass's Net, S=1,0 m2)․
1 biomass is calculated on nomogrammes ( Chislenko, 1968) and equivalent to wet weight
2 ratio of mean abundance (or biomass) between M. longiseta nov.sp. and total Halocyprididae
TABLE 1. Meristic characteristics of Metaconchoecia longiseta nov. sp.
| Characteristic | Range | Mean and number specimens |
|---|---|---|
| Length (mm): male | 1,301,55 | 1,47 (n=21) |
| female | 1,401,60 | 1,49 (n=46) |
| Height (%): male | 45,3351,05 | 47,63 (n=21) |
| female | 47,3053,79 | 50,0 9 (n=45) |
| Width (%): male | 45,1650,72 | 47,53 (n=21) |
| female | 38,9350,34 | 42,0 6 (n=43) |
| LAG (%): both sexes | 12,0014,49 | 12,77 (n=13) |
| Penis (% length of carapace) | 27,5935,81 | 27,71 (n=9) |
TABLE 2. The comparison of characteristics among M. skogsbergi, M. longiseta nov. sp. and M. alta nov. sp.
| Characteristic | M. skogsbergi | M. longiseta nov. sp. | M. alta nov. sp. |
|---|---|---|---|
| Carapace (both sexes): | |||
| height | considerably more in posterior half | considerably more in posterior half | slightly more in posterior half |
| right asymmetrical gland | opens above posterodorsal corner | opens at (or almost) posterodorsal corner | opens at (or almost) posterodorsal corner |
| Male. Frontal organ: | |||
| capitulum | wide distally | narrow distally | wide distally |
| shaft | barely or not reachs of boundary between 2nd and 3rd segments of 1st antenna | extends beyond boundary between 2nd and 3rd segments of 1st antenna | extends beyond boundary between 2nd and 3rd segments of 1st antenna |
| First antenna: | |||
| aseta | shorter than limb | shorter than limb | longer than limb |
| bseta | shorter than dseta | longer than dseta | longer than dseta |
| cseta | thin and long | thin and long | thick and short |
| Endopodite of second antenna: |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Metaconchoecia
| Chavtur, Vladimir Grygorjevich 2003 |
Metaconchoecia
| Chavtur 1991: 46 |
Metaconchoecia skogsbergii:
| Chavtur 1976: 105 |