Macrogomphus albardae Selys, 1878

Kosterin, Oleg E., 2019, New synonyms and a new subspecies of Macrogomphus Selys, 1858 (Odonata: Gomphidae) from continental south-east Asia, Zootaxa 4615 (1), pp. 57-90 : 59

Macrogomphus albardae Selys, 1878 ,?bona species

Figs. 2–8

References to records from mainland Asia (excluding the Malay Peninsula)

Macrogomphus rivularis n. sp. — Förster (1914): 80 –82 (original description).

Macrogomphus borikhanensis sp. nov. — Fraser (1933): 136 –139, fig. 8a, c (original description; drawings of anal appendages).

borikhanensis (Macrogomphus) — Kimmins (1966): 183 (lectotype designation).

borikhanensis Fraser (Macrogomphus) — Kimmins (1969): 292 (type specimens listed).

Macrogomphus borikhanensis — Asahina (1981): 8 (report for Thailand, Chanthaburi Province).

Macrogomphus guilinensis Chao , sp. nov. — Chao 1982: 11–13, figs. 1–6 (original description; drawings of anal appendages, secondary genitalia and synthorax).

Macrogomphus borikhanensis — Asahina (1986): 7 –9, fig. 1–6 (description and figure of the Chanthaburi specimen).

Macrogomphus guilinensis Chao—Chao (1990) : 156–157, fig. 5–6.1 (original description reproduced and commented).

Macrogomphus sp.— Yokoi (1999): 4, fig. 9 (report for Laos, 30 km S of Vientiane).

Macrogomphus species, apparently borikhanensis — Pinratana (2000): 155 (report for Thailand, Lop Buri Province).

Macrogomphus matsukii Asahina, 1986 — Yokoi (2001): 26 (report for southern Laos, Sekong).

rivularis Förster (Macrogomphus) , 1914:80— Garrison et al. (2003): 84 (holotype listed).

Macrogomphus guilinensis Chao, 1982 — Wilson (2005): 136, fig. 17 (redescription of male and the first description of female from Nonggang, Guangxi Zhuang Autonomous Region, drawings of male appendages and female head, synthorax, abdomen and vulvar scale).

Macrogomphus matsukii Asahina, 1986 — Kosterin et al. (2011): 14 –15 (a record for Thailand, Nakhon Ratchasima Province, Nong Bun Nak village).

? Macrogomphus guilinensis Chao, 1982 — Karube (2014): 156 –157, fig. 13 (report for Vietnam, Vĩnh Phúc Province, Tam Đảo Mountain; drawings of the anal appendages).

Macrogomphus borikhanensis Fraser, 1933 — Yokoi & Souphanthong (2014): 16 (mentioned for Laos with reference to Fraser 1933)

Macrogomphus matsukii Asahina, 1986 — Yokoi & Souphanthong (2014): 16, 89 (mentioned for Laos with reference to Yokoi 2001 and three localities in Vientiane Province and Prefecture and Attapu Province indicated on the map)

Macrogomphus matsukii Asahina, 1986 — Kosterin (2017): 29, fig. 28 (a record for Cambodia, Stung Treng Province, 2 km SW of Kampong Preah village, photo of a female habitus).

Macrogomphus guilinensis — Kompier (2019) (reports for Bắc Kạn Province including Ba Bể National Park, photos of male and female habitus, male appendages and female tip of abdomen).

Macrogomphus matsukii — Kompier (2019) (a photo of a male in nature from southern Vietnam, Cát Tiên National Park)

Figures

FIGURE 2. Details of some male specimens of Macrogomphus albardae kept in RMNH (courtesy of Jan van Tol): a–d—the male syntype (the labels ‘Holotype’ are misleading); e—“W. Borneo, 0 m. / Singkawang / Penanring / L. Coomans de Ruiter / 24. II. 1932”; f—“S. Sumatra / Terbanggi-Hilir / nr. Menggala / Lampang distr. / 18. VIII. 1936 / M. Bartels”; a—general habitus; b, e–f—S10 and anal appendages, dorsal view; c—head and thorax, dorsolateral view; d—labels of the male syntype. Not to scale.

FIGURE 3. Anal appendages (a–h) and secondary genitalia (j–k) of Macrogomphus albardae as illustrated in literature: aafter Lieftinck (1935: fig. 9) (as M. parallelogramma albardae); b—after Orr (2005: p. 64) (as M. parallelogramma albardae); c—after Fraser (1933: fig. 8a, c) (as M. borikhanensis); d—after Asahina (1986: fig. 8–9) (as? M. phalantus): e, j—after Asahina (1986: fig. 4–6) (as M. borikhanensis); f, i—after Chao (1982: fig. 1, 3, 6) (as M. guilinensis); g—after Wilson (2005: fig. 17 d, e) (as M. guilinensis); h—after Karube (2014: fig. 13) (as M. guilinensis). Not to scale; j mirrored. Asterisk indicates the proximal inner swelling of the cercus.

FIGURE 4. The dashed (a) and intermediate (b) morphs of males of Macrogomphus albardae in nature in peninsular Thailand, Koh Samui Island, Baan Lipa Noi, Wanom Waterfall: a—10 iii 2012; b—13 vii 2009. Courtesy of Leslie Day.

FIGURE 5. A small male of Macrogomphus albardae of the ringed morph, collected by the author on 21 viii 2018 in Cambodia, Preah Vihear Province, Srae Veal village env.: a—habitus in live colours, dorsal view; b—ditto, lateral view (not to scale); canal appendages, dorsal view; d—ditto, lateral view; e—secondary genitalia, lateral view; f—posterior hamulus, anteriolateral view. Scale bars: upper 2 mm; lower 0.5 mm.

FIGURE 6. Holotype of Macrogomphus rivularis Förster, 1914 (a–b) and its labels (c) (UMMZ): a—dorsal view; b—lateral view. Courtesy of Erica Tucker.

FIGURE 7. Details of the abdomen of the holotype of Macrogomphus rivularis Förster, 1914 (UMMZ): a—end of abdomen, dorsal view; b—ditto, lateral view; c—ditto, ventral view; d—base of abdomen with secondary genitalia, lateral view. Not to scale. Courtesy of Erica Tucker.

FIGURE 8. Holotype of Macrogomphus borikhanensis Fraser, 1933 and its labels (BMNH): a—dorsal view; b—lateral view. Not to scale. Courtesy of Benjamin Price.