Prolycosides aussereri ( Keyserling, 1877 ) Paredes-Munguía & Brescovit & Teixeira, 2024

Paredes-Munguía, Williams, Brescovit, Antonio D. & Teixeira, Renato A., 2024, Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species, Zootaxa 5414 (1), pp. 1-83 : 59-66

Prolycosides aussereri ( Keyserling, 1877) comb. nov.

Figs 52‒56 , 67

Tarentula aussereri Keyserling, 1877: 657 , pl. 7, fig. 33. Female holotype from St. Fé de Bogota, Colombia, deposited in the BMNH-1890/999, examined by photographs ( Fig. 56 ).

Lycosa aussereri : Banks 1901: 223.

Arctosa aussereri : Roewer 1955: 230.

Lycosa proletaria Tullgren, 1905: 65 , pl. 8, fig. 31. Female holotype from Tatarenda, ca. 19°6'23.24"S 63°30'51.22"W, Chuquisaca, Bolivia, 1901-1902, E. Nordeskiöld leg., deposited in NRS, Sweden, 18, examined. Mello-Leitão 1941a: 134, fig. 28. Syn. nov.

Avicosa proletaria : Roewer 1955: 236.

Schizocosa proletaria : Chamberlin & Ivie 1942: 38.

Tarentula workmani Strand, 1909: 277 . Female holotype from Paraguay, ix.1880, should be in National Museum of Ireland, Dublin, Ireland, lost). Syn nov.

Hygrolycosa workmani : Roewer 1955: 261.

Arctosella workmani : Roewer 1960: 671.

Lycosa planithoracis Mello-Leitão, 1938: 98 , figs 12–13. Female holotype from Chacras de Coria , ca. 32°58'51.27"S 68°52'36.84"W, Mendoza, Argentina, Don P. Jorgensen leg., deposited in MLP 14059, examined. Syn. nov.

Hogna planithorax : Roewer 1955: 256. (unacceptable emendation).

Lycosa variolosa Mello-Leitão, 1941a: 134 , fig. 32, pl. VI, fig. 26. Female holotype from Villa Nueva, ca. 32°26'41.65"S 63°15'16.17"W, Córdoba, Argentina, A. Parko leg., deposited in MLP 14702, examined. Syn. nov.

Lycorma variculosa : Roewer, 1955: 267 (Transferred from Lycosa , lapsus).

Lycosa melanostoma Mello-Leitão, 1941b: 208 , fig. 7. Female holotype from Reconquista, ca. 29°8'5.20"S 59°38'32.11"W, Santa Fé, Argentina, M. Birabén leg., deposited in MLP 15076, examined. Syn. nov.

Megarctosa melanostoma : Roewer 1955: 278.

Lycosa huachoi Mello-Leitão, 1942a: 431 , fig. 5. Female holotype from Huacho, 11°6′S 77°36′W, Peru, deposited in MNRJ 02001, examined, and subsequently destroyed in the 2018 fire. Syn. nov.

Ocyale huachoi : Roewer 1955: 281.

Lycosa abalosi Mello-Leitão, 1942b: 391 , fig. 4. Female holotype from Colonia Dora, ca. 28°36'25.32"S 62°57'9.57"W, Santiago del Estero, Argentina, deposited in MLP 15310, examined. Syn. nov.

Sosilaus abalosi : Roewer 1955: 296.

Pirata abalosi : World Spider Catalog 2023

Lycosa soukupi Mello-Leitão, 1942a: 431 , fig. 4 (female holotype from Huacho, 11°6′S 77°36′W, Peru, deposited in MNRJ 01364, examined, and subsequently destroyed in the 2018 fire. Syn. nov.

Sosilaus soukupi : Roewer 1955: 296.

Pirata soukupi : World Spider Catalog 2023

Other material examined. CUBA: Piñar del Rio: 1♂, La Bajada, Peninsula de Guanahacabibes , ca. 21°54'17.19"N 84°23'46.51"W, 12.v.2002, A. Sanchez leg. ( IBSP 169614 ) . Sabaera: 1♀, Maisí, Guantánamo , ca. 20°13'32.99"N 74°9'12.08"W, 12.iv.1998, A. Sanchez leg. ( IBSP 169619 ) , 1♀, ditto, 4.iv.1998, A. Sanchez leg. ( IBSP 169610 ) , Santiago de Cuba: 2♂, La Puya, III Frente ,ca. 20°8'45.38"N 76°17'38.79"W, 19.iii.2000, A. Sanchez-Ruiz leg. ( IBSP 233858 ) . VENEZUELA: Maracay: 1♀, Radio Sertaneja , ca. 10°15'32.50"N 67°38'27.46"W, 454 m, 16.iv.1984, W. Manya leg. ( MUSM-ENT 500166 ) . COLOMBIA: Cundinamarca: 1♀, Cáqueza, confluence of Cáqueza and Rio Negro rivers, ca. 4°23'53.50"N 73°54'20.33"W, 1520 m, 4.vi.2002, A. Rodriguez leg. (ICN-Ar 1867) ; 1♀, Sasaima , 19.x.1997, R.Casallas leg. ( MLS 302 ) ; 1♀, Fusagasugá ,ca. 4°20'42.55"N 74°21'42.56"W, 14.vi.1905, R. Guevara leg. ( MLS 291 ) . Cauca: 1♀, El Tambo, ca. 2°27'07.59"N 76°48'41.21"W, 600 m, 29.xii.1998, R. Casallas leg. ( MLS 436 ) . PERU: Tumbes: 1♂, Tumbes, Corrales , ca. 3°34'S 80°28'W, 5 m, ii.1983, P. Vinces & P. Castillo leg. ( MUSM-ENT 500151 ) ; 1♀, Palmas, Rio Queiroz , ca. 3°34'00.22"S 80°12'50.71"W, 600 m, 8.x.1955, F. Blancas leg. ( MUSM-ENT) . Piura: 1♀, near Huancabamba, Valle N. Pucara , 5°56.15'S 79°16.19'W, 1058 m, 20.viii.2006, G. Binford et al. leg. ( MUSM-ENT) ; 1♀, Ayabaca , ca. 4°38'S 79°43'W, 2715 m, 24.ix.2019, F. Blancas leg. ( MUSM-ENT) ; 1♂, Chocan, Ayabaca , ca. 4°41'31.23"S 80°34'04.42"W, 114 m, 2.x.1955, F. Blancas leg. ( MUSM-ENT 500131 ) . Amazonas: 1♀, Rodriguez de Mendoza, Limabamba , ca. 6°29'52.43"S 77°29'55.71"W, 1674 m, 6.xi.1986, R. Fernandez leg. ( MUSM-ENT 500133 ) . Cajamarca: 2♂, Celendín, road to Llangoat , 6°42'51"S 77°51'55.9"W, 2048 m, 7.xii.2004, W. Paredes leg. ( MUSM-ENT) ; 1♂, La Victoria , ca. 6°57'37.85"S 78°06'53.24"W, 2638 m, 23.vi.1986, H. Ortega leg. ( MUSM-ENT 500159 ) ; 1♂, Baños del Inca , ca. 7°10'S 78°28'W, 2659 m, 6.viii.1984, I. Bohorquez leg. ( MUSM-ENT 500144 ) ; 1♂, San Andres de Cutervo , ca. 6°22'S 78°49'W, 2866 m, 16.iii.1989, D. Silva leg. ( MUSM-ENT) ; 2♂, Rio Mashcón , ca. 6°12'16.65"S 78°38'16.03"W, 240 m, 4.x.1965, CC & BB leg. ( MUSM-ENT) ; 1♀, Hacienda Sunchubamba , 7°29'S 78°23'W, 2657 m, 2.xi.2009, D. Silva leg. ( MUSM-ENT) ; 3♀, Cumbemayo , ca. 7°11'26.07"S 78°34'43.23"W, 3592 m, 27.xi.2004, W. Paredes leg. ( MUSM-ENT 500154 ) . Lambayeque: 1♂, Motupe , ca. 6°45'07.24"S 79°51'07.68"W, 24 m, 2.ix.2009 ( MUSM-ENT) ; 1♀, Chiclayo , ca. 6°47'24.35"S 79°51'30.35"W, 21 m, 22.vii.2006, A. Albujar leg. ( MUSM-ENT) ; 1♀, Motupe, Distrito de Olmos , ca. 5°59'15.81"S 79°44'42.91"W, 142 m, 21.ii.1955, Vellard leg. ( MUSM-ENT 500129 ) ; 1♀, Lambayeque, Santa Maria dos Vales de Chiclayo , ca. 6°45'47.00"S 79°50'12"W, 27 m, 2.viii.2006, A. Albujar leg. ( MUSM-ENT) . La Libertad: 3♀, Chicama, Valle de Chicama , ca. 7°52'59.94"S 79°13'00.04"W, 61 m, 1.ix.1978, P. Hocking & G. Martinez leg. ( MUSM-ENT 500138 ) ; 3♂ 1♀, Pataz, Yalen , ca. 7°45'32.08"S 77°33'37.13"W, 2850 m, 26.iii.1988, D. Silva leg. ( MUSM-ENT 500135 ) ; 1♀, Trujillo, Victor Larco Herrera, Santiago de Huamán , ca. 8°08'18.98"S 79°02'23.28"W, 10 m, iii.1983, Sinuas leg. ( MUSM-ENT) . Ucayali: 1♀, Tipishca, Tamaya , ca. 8°27'56.84"S 74°14'36.38"W, 149 m, 13–19.vi.1958, Koepcke & Koepcke leg. ( MUSM-ENT 500165 ) ; 1♂ 1♀, Ivita, near to Pucallpa , ca. 8°38'S 74°58'W, 196 m, 23–24.vii.1986, D. Silva leg. ( MUSM-ENT 500145 ) ; 1♀, ditto, 4.x.2002, R. Ramirez leg. ( MUSM-ENT 500158 ) . Huánuco: 1♀, Rio Yuyapichis, Panguana Biological Station , 9º37'S 74º56'W, 236 m, 26.ii–24.xii.1983, M. Verhaghi leg. ( IBSP 23680 ) ; 1♂, Huacaybamba, Road Huambu-Huanuco, near Hui Cuibambu , ca. 9°31'36.18"S 76°46'02.24"W, 3511 m, x.1984, R. Tejada leg. ( MUSM-ENT) ; 1♂ 1♀, Tingo Maria , ca. 9°17'39.51"S 75°59'14.64"W, 705 m, 14–16.vii.1948 ( MUSM-ENT) . Pasco: 1♂, Oxapampa , ca. 10°34'30.09"S 75°24'00.04"W, 1817 m, 22.vi.1986, D. Silva leg. ( MUSM-ENT 500150 ) . Lima: 1♀, San Vicente de Cañete, Valle de Mala , 13º5'S 76º24'W, 19 m, 18.ix.1997, M.A.B. Pino leg. ( IBSP 14893 ) ; 4♂ 1♀, ditto ( IBSP 14899 ) ; 1♀, ditto, 9.xi.1997 ( IBSP 14895 ) ; 1♂, ditto, 7.xi.1997 ( IBSP 14906 ) ; 3♂, ditto ( IBSP 14913 ) ; 1♂, ditto, 19.x.1997 ( IBSP 14915 ) ; 2♂, ditto ( IBSP 14919 ) ; 1♂ 2♀, ditto, 7.ix.1997 ( IBSP 14921 ) ; 1♀, Chorillos, Los Pantanos de Villa , ca. 12°12'22.00"S 76°59'20.00"W, 2 m, 19.x.2003, W. Paredes leg. ( MUSM-ENT 500142 ) ; 1♂, ditto, 20.viii.2006 ( MUSM-ENT) ; 1♀, Road to Canta, Km 40, Trapiche , Recreo La Retama , 11°43'20.80"S 76°57'46.7"W, 546 m, 23.vi.2006, W. Paredes leg. ( MUSM-ENT) ; 1♀, Huacho, Rio Huacho , 11°04'39.16"S 77°36'03.33"W, 47 m, 21.x.2006, W. Paredes leg. ( MUSM-ENT) ; 1♂, Hurin, La Tablada, Rio Lurin , ca. 12°12'27.83"S 76°52'12.27"W, 84 m, 12.vii.1988, G. Soza leg. ( MUSM-ENT) ; 1♀, Cañete, Rio Cañete , ca. 13°07'29.65"S 76°21'44.05"W, 49 m, 15.ii.1996, S. Cordova & J. Duarez leg. ( MUSM-ENT 500152 ) ; 1♀, San Antonio de Chaclla, Jicamarca , ca. 11°54'23.68"S 76°42'12.53"W, 450 m, 1.ix.1968, A. Hoempper leg. ( MUSM-ENT 500137 ) ; 1♀, La Molina , 12°5'S 76°57'W, 236 m, 1.i.2006, A. Aparicio leg. ( MUSM-ENT 500126 ) ; 1♀, ditto, 15.i.2006, A. Aparicio leg. ( MUSM-ENT 500127 ) . Ica: 2♀, Huacachina , ca. 14°05'16.27"S 75°45'40.72"W, 401 m, 4.x.1985, F. Blancas leg. ( MUSM-ENT) . Cusco: 1♀, Santa Maria, Quillabamba , ca. 12°52'S 72°42'W, 1070 m, ix.2002, G. Binford et al. leg. ( MUSM-ENT) . Puno: 1♀, Puno, Pukará , ca. 15°02'30.51"S 70°21'48.11"W, 3867 m, 6.viii.2004, G. Binford et al. leg. ( MUSM-ENT) . Arequipa: 1♀, 7 km E. Cháparra, ca. 15°41'S 73°49'W, 1450 m, 15.ix.1988, R. Tejada leg. ( MUSM-ENT 500134 ) ; 1♀, ditto, 1496 m, 19.ix.1988, R. Tejada leg. ( MUSM-ENT) . BOLIVIA: Cochabamba: 1♀, Near Monte Puncu , ca. 17°35'13.82"S 65°18'23.36"W, 2700 m, 22.x.1983, A. Roig leg. ( MACN) ; 1♀, ditto ( MACN) . BRASIL: Pernambuco: 1♂, Brejão, Fazenda Santa Rita, Vila Santa Rita , ca. 7°33'19.85"S 35°0'5.91"W, v–x.2002, S.D. Bella leg., ( IBSP 41085 ) . Acre: 2♂ 1♀, Rio Branco, Reserva Extrativista Catuaba , ca. 10°48'29.08"S 68°43'22.02"W, 262 m, 9.iv.1996, Equipe IBSP & SMNK leg. ( IBSP 15953 ) . Amapá: 1♂ 2♀, Laranjal do Jari, Vila Cachoeira de Santo Antônio , ca. 0°49'54.59"S 52°30'30.83"W, 15 m, 18–24.ii.2003, J.A.P. Barreiros leg. ( IBSP 53787 ) ; 1♂, Serra do Navio , 0°58'N 52°02'W, 110 m, 16–17.vi.1966, Galiano leg. ( MACN) ; 1♂ ditto, ( MACN) . Amazonas: 1♀, Coari, Rio Urucu, Porto Urucu , ca. 4°50'55.47"S 65°20'28.17"W, 61 m, 13.vii.2003, A.B. Bonaldo leg. ( IBSP 53789 ) ; 1♀, Benjamin Constant , ca. 4°23'33.29"S 70°01'19.25"W, 79 m, ix.1962, K. Lenko leg. ( MZUSP 20517 ) . Pará: 1♀, Joanes, Ilha de Marajó , ca. 0°52'42.65"S 48°30'33.22"W, 12 m, 7.x.2002, A.B. Bonaldo leg. ( IBSP 53799 ) ; 2♀, Santarém, Fátima de Urucurituba , ca. 2°26'28.78"S 54°48'46.64"W, 73 m, 24.i.1994, A.D. Brescovit leg. ( MCN 25021 ) . Bahia: 1♀, Xique-Xique , ca. 10°49'16.44"S 42°43'01.57"W, 405 m, iii.1966 ( IBSP 1984 ) ; 1♀, Iraquara, Chapada Diamantina , ca. 12°14'44.11"S 41°37'25.99"W, 691 m, 15.xii.1998, L.S. Rocha leg. ( IBSP 53876 ) . Mato Grosso do Sul: 3♀, Serra da Bodoquena , ca. 21°05'07.82"S 56°39'40.16"W, 616 m, 20–22.xi.2001, J. Raizer leg. ( IBSP 33023 ) . Rio de Janeiro: 1♂ 1♀, Maricá, Praia de Bambuí , ca. 22°56'44.27"S 42°45'25.71"W, 2 m, 9.v.1977, C.J. Becker leg. ( MCN 6179 ) ; 1♀, São João da Barra, Atafona , ca. 21°38'45.77"S 41°01'09.92"W, 4 m, 07.viii.1963, Exp. Dpto. Zool. leg. ( MZUSP 20481 ) . São Paulo: 1♀, Barão de Itapetininga , ca. 23°35'28.72"S 48°02'44.27"W, 663 m, 2.vii.1964 ( IBSP 1888 ) ; 1♀, São Paulo, 779 m, ( IBSP 1948 ) ; 1♀, São Paulo, 23º32'51"S 46º38'10"W, 789 m, 17.vii.1969, D. Martins leg. ( IBSP 2283 ) ; 1♀, Araçoiaba da Serra , ca. 23°30'43.03"S 47°36'30.44"W, 625 m, ii.1987, R. Klink leg. ( IBSP 4552 ) ; 1♀, Osasco , 23º31'19"S 46º48'07.77"W, 720 m, iii.1990, Equipe IBSP leg. ( IBSP 5041 ) ; 1♂, Natividade da Serra , ca. 23°22'45.81"S 45°26'37.54"W, 746 m, 23–26.vi.2001, M. Calleffo leg. ( IBSP 29171 ) ; 1♀, Guarulhos , ca. 23°27'17.58"S 46°31'48.61"W, 767 m, 01.iv.2003, B. Imporato leg. ( IBSP 37144 ) ; 1♂, São Paulo, ca. 23°32'38.07"S 46°38'30.61"W, 790 m, xi.1981, M.A. Klaussner leg. ( IBSP 53807 ) ; 1♀, Itapevi , 23º32'51.80"S 46º56'24.55"W, 742m, 08.iv.1999, R. Sales leg. ( IBSP 23650 ) . 1♂ 1♀, Presidente Prudente, Rosana, Usina Hidrelétrica Porto Primavera , ca. 22°06'47.06"S 51°27'53.42"W, 394 m, 11.xii.1998 – i.1999, Eq. IBSP leg. ( IBSP 53808 ) ; 1♀, ditto, Eq. IBSP leg. ( IBSP 53836 ) ; 2♀, Rosana , ca. 22°09'07.56"S 51°24'33.29"W, 424 m, 21.ii.2001, Usina Hidrelétrica Sérgio Motta, R.P. Indicatti & D.F. Candiani leg. ( IBSP 53817 ) ; 7♀, ditto, ( IBSP 53818 ) ; 1♀, ditto ( IBSP 53838 ) ; 4♂ 6♀, ditto, 11.xii.1998 – i.1999 ( IBSP 53846 ) ; 1♀, ditto, ( IBSP 53850 ) ; 2♂ 2♀, Rosana , 22º34'47"S 53º03'33"W, xii.1986, Equipe IBSP leg. ( IBSP 4499 ) . Rio Grande do Sul: 1♂, Rio Pardo , ca. 29°58'54.75"S 52°23'39.11"W, 17 m, 20.ii.1967, A.A. Lise leg. ( IBSP 53791 ) ; 1♀, Herval, Santa Eliza , ca. 32°01'38.49"S 53°40'47.37"W, 101 m, 28.vii.1973, J. Sbruzzi leg. ( MCN 3565 ) ; 1♀, Salto do Jacuí, Itaúba , ca. 29°04'58.58"S 53°13'12.95"W, 289 m, 11.iv.1978, A.A. Lise leg. ( MCN 7933 ) ; 1♀, Santa Maria, Camobi , ca. 29°42'58.58"S 53°43'36.55"W, 112 m, i– vi.1982, M. Ceolin leg. ( MCN 14319 ) ; 1♀, Eldorado do Sul, ca. 30°00'15.96"S 51°18'10.79"W, 7 m, 30.ii.1996, G.S. Carvalho & R. Silva Jr. Leg. ( MCTP 12585 ) ; 1♂, ditto, 11–17.i.1997 ( MCTP 12604 ) ; 9♀, Reserva Biológica de São Donato, São Borja , ca. 29°02'49.09"S 56°03'05.09"W, 93 m, 23.vii.2012, M. Machado leg. ( MCTP 36211– 36214 ) ; 1♀, ditto, 11.vii.2012 ( MCTP 36226 ) . PARAGUAY: Central: 1♀, Villeta , ca. 25°30'25.26"S 57°33'32.18"W, 92 m, 30.viii.1984, J. Kochalka leg. ( IBNP) ; 1♀, ditto, ( IBNP) ; 1♂, ditto, 24.vii.1982 ( IBNP) ; 1♂, San Lorenzo , ca. 25°20'27.34"S 57°30'44.39"W, 118 m, 1–10.x.1985, J. Kochalka leg. ( IBNP) ; 1♂ 2♀, Asunción , 14.ix.1984, J. Kochalka leg. ( IBNP) ; 1♀ ditto, 14.ix.1984 ( IBNP) ; 1♀ ditto, 14.ix.1984 ( IBNP) . Boquerón: 4♂ 8♀, Estancia La Gama , ca. 23°04'32.66"S 61°28'11.82"W, 191 m, 19–20.x.1990, J. Kochalka leg. ( IBNP) ; 1♂, ditto, ( IBNP) .

Itapúa: 1♂, Isla Yacyretá , ca. 27°26'00.35"S 56°45'55.70"W, 73 m, 12–15.ix.1992, J. Kochalka leg. ( IBNP) ; 1♀, Near Arroyo Guapó, General Artigas, San Miguel Potrero , Coronel Bogado , ca. 27°10'14.89"S 56°13'03.97"W, 105 m, 29–30.ii.1996, J. Kochalka leg. ( IBNP) . 1♂, Isla Yacyretá, near Estrada Melgarejo , 27°25′48"S 56°46′12"W, 70 m, 12–15.ix.1992, J. Kochalka leg. ( IBNP) ; 1♀, ditto, 25.iv.1993, D. Torres leg. ( IBNP) . Paraguarí: 3♂, Presidente Hayes, 25 Leguas , ca. 22°47'50.74"S 59°36'34.56"W, 121 m, 11–12.vii.1983, J. Kochalka leg. ( IBNP) ; 3♂, 25 Leguas (currently Teniente Primero Manuel Irala Fernández ), 22°48′50.4″S 59°35′9.6″W, 120 m, 11–12.vii.1963, J. Kochalka leg. ( IBNP) ; 1♂, Ruta Trans-Chaco , km 20, ca. 25°10'41.50"S 57°33'07.91"W, 60 m, 25.viii.1981, J. Kochalka leg. ( IBNP) . ARGENTINA: Salta: 1♀, Tartagal , ca. 22°31'00.34"S 63°48'20.19"W, ii.1963, Apostoli leg. ( MACN) ; 1♀, Rio Santa Maria , ca. 26°01'28.45"S 65°52'05.24"W, vii.1947, A.G. Giai leg. ( MACN 2275 ) ; 1♀, Lorenzo , ca. 24°43'57.72"S 65°29'12.85"W, 6.vi.1927, Zotta leg. ( MACN) . Buenos Aires: 4♀, Berazategui , ca. 34°45'45.50"S 58°12'37.49"W, 16.vi.1940, F. Monrós leg. ( MACN) . Corrientes: 1♀, Itati , ca. 27°16'22.07"S 58°14'33.23"W, 26.viii.1963, M.E. Galiano leg. ( MACN) . Santiago Del Estero: 1♀, Beltran , ca. 27°49'46.28"S 64°03'39.75"W, vi.1940, Birabén leg. ( MLP) . Tucuman: 1♀, Güemes , ca. 27°18'50.65"S 65°19'51.43"W, 26.x.1998, ( MACN) . 4♀, Burruyacú , ca. 26°30'02.96"S 64°44'30.95"W, xi.1960, Birabén leg. ( MACN) ; 1♀, ditto, xi.1960, M. Birabén leg. ( MACN) . Entre Rios: 1♀, Rincón de Nogoyá , ca. 32°46'45.70"S 59°54'36.08"W, 27.ii.1982, A. Roig leg. ( MACN) ; 1♀, Parque Nacional El Palmar , 31°51'11"S 58°19'21"W, 18.iv.1981, P. Goloboff leg. ( MACN) .

Formosa: 1♀, Parque Nacional Pilcomayo , ca. 25°04'01.10"S 58°07'48.16"W, ix.1990, M.J. Ramírez leg. ( MACN) .

Chaco: 1♀, Resistencia , ca. 27°25'16.11"S 58°59'17.68"W, xii.1976, Aguillera leg. ( MACN) ; 1♀, ditto, Denier leg. ( MACN 82 ) .

Remarks: Prolycosides was erected by Mello-Leitão (1942b) from a single female based on the following diagnostic characters: sparse spinulation on tibiae I and II, atypical compared to the Lycosa pattern, and five small retromargin equal teeth lower border as well as the broad, smooth, and rectangular atrium on epigynum. Although spinulation patterns are not always useful for distinguishing between genera, as demonstrated in other families (e.g., Hippa & Lehtinen 1983 [ Lycosidae ]; Ramírez 2003 [ Anyphaenidae ]; Miller et al. 2012 [ Eresidae ]) Mello-Leitão succeeded in separating Prolycosides from other genera based on this evidence. After examination of the type specimen and paired with males, we added diagnostic characters for the genus based on the males: embolus so long that it bends at the tegular lobe and absence of basal apophysis at the median apophysis.

Different from Hogna , Prolycosides species have a straight prolateral arm of the median apophysis, which is curved in Hogna, As mentioned, the tip of the embolus rests on the tegular lobe but is not wrapped by it as in Pavocosa . In Prolycosides females the median septum is twice as long as the transversal septum ( Fig. 53b ), and not thrice as it usually is in Pavocosa . By contrast the median septum in Hogna is quite the same length as the transversal septum, with different degrees of dilatation on median septum ( Dondale & Redner 1990). The atrium in Prolycosides is smooth and wider ( Fig. 55b ) than in Hogna , and not narrow or striated as in Pavocosa ( Fig. 55a ).

Diagnosis. The males of Prolycosides have a very large embolus with a bent tip at the tegular lobe ( Fig. 54e ) and lack a basal protuberance on the median apophysis ( Fig. 54b ). Females differ from other genera in having a broad, rectangular atrium ( Fig. 53b ) and an ovoid spermatheca head ( Fig. 53e ).

Description. Male (MUSM-ENT 500135). Carapace brown, covered by short black bristles except at longitudinal median band and submarginal bands, both yellow ( Fig. 53a ), broad at the front, tapering towards the back. Sparse black bristles between the ocular area and the thoracic groove. Ocular area black, covered with white bristles, cephalic portion of the carapace covered by sparse, large white bristles ( Fig. 53h ). Sternum longer than wide, uniformly light brown, covered by short black bristles, which are sparse in the marginal area ( Fig. 53d ). First row of eyes slightly procurved, shorter than the second row; AME slightly larger than the ALE; PME more than a radius apart. Endites yellow, teardrop-shaped and convergent; labium square, darker than the endites; both endites and labium yellowish in their distal portion. Chelicerae brown, frontally covered with sparse black setae and dense white setae in all its extension; striae file on its frontal part; chillum membranous, divided with the mesal portion sclerotized; condyle large and clearer than the chelicerae. Legs: All segments uniformly yellow except on tarsi and metatarsi I and II, which are darker. Opisthosoma entirely covered by alternated tiny, black, and white bristles, the background dark olive; cardiac mark dark, flanked by yellow longitudinal bands ( Fig. 53a ). Venter uniformly pale yellow from epigastric furrow to spinnerets; anterior spinnerets wider than posterior spinnerets, both cylindrical ( Fig. 53d ).

Pedipalp ( Figs 52 , 54 ), tibia cylindrical. Cymbium piriform (Fig, 52b), subtegulum lenticular ( Fig. 52h ).Tegulum with a curved, C-shaped sperm duct. Terminal apophysis sclerotized, laminar, sickle-shaped, and distally sharp ( Figs 52e , 54a ), subterminal apophysis membranous, shorter than terminal apophysis ( Figs 52g , 54b ). Embolus thin, with tip curved and embraced by the lamellar part of tegular lobe ( Figs 52e, g , 54b ). Median apophysis triangular and with a sharp tip ( Figs 52d , 54a ); ventral projection (VP) short and slightly curved ventrally ( Figs 52a , 54a ).

Leg formula IV>I>II>III. Spination pattern: femur I p 0-0-2 d 1-1-1 r 1-1d-1d, II p 1d-1d d 1-1-1 r 1-1-1, III-IV p 1d-1d d 1-1-1 r 1d-1d; patella I-IV p 1 d 1bristle-1bristle ap r 1; tibia I-II p d1-d1 d 1bristle-1 v 2-2-2ap r 1-1d, III-IV p 1d-1d d r1-1 r 1-1 v 2-2-2ap; metatarsus I-II p 1d-1d-2 r 1d-1d-2 d 2bristle-2bristle v 2-2-1ap, III p 1d-1d-2 d 1bistle-bristle-0 r 1d-1-2 v 2-2-1ap, IV p 1d-1d-2 d 1bistle-1bristle-0 r 1d-1d-2 v 2-1-2-1ap.

Measurements: TL 15.06, CL 6.78, CCW 2.83, CTW 5.22, CCH 2.22, CTH 2.22. Eyes: AME 0.29, ALE 0.24, PME 0.60, PLE 0.45, interdistances: AME-AME 0.17; AME-ALE 0.06; PME-PME 0.40; PLE-PLE 1.21. POQ long 0.95, POQ posterior width 1.54, POQ anterior width 0.98. Chelicerae: length 2.55. Opisthosoma: length 6.78, width 4.66. Legs: femur + patella + tibia + metatarsus + tarsus = total length: pedipalp 2.55 + 1.22 + 1.39 + - + 1.83 = 6.99, I 5.94 + 2.78 + 5.22 + 5.44 + 3.00 = 22.38, II 6.17 + 2.61 + 4.78 + 5.00 + 2.89 = 21.45, III 5.00 + 2.22 + 4.22 + 5.67 + 2.78 = 19.89, IV 6.33 + 2.56 + 5.44 + 7.78 + 3.33 = 25.44.

Female (MUSM-ENT 500135). General pattern as in male. First row of eyes straight, shorter than the second row; AME slightly larger than the ALE; PME separated by a length same as their radii. Chelicerae with three promarginal teeth, the median the largest; retromarginal with three teeth, the basal the shortest. Chelicerae is darker brown than in males; chillum as in males; boss larger compared with males. Sternum, endites and labium yellow ( Fig. 53f ). Legs and opisthosoma as in males ( Fig. 53i ).

Genitalia ( Figs 53b, e , 55 ), LS and TS forming a T-shaped plate ( Fig. 53b ). LS wider than TS, atrium smooth and wide ( Fig. 55a ). CO at the border of TS ( Fig. 55c ). Hoods parallel and triangular ( Figs 53b , 55a ). HS kidneyshaped, SS narrow; BS bulbous, VC absent ( Figs 53e ).

Leg formula IV>I>II>III. Spination pattern: femur I p 0-0-2 d 1-1-1, II p 0-d1-d1 d 1-1-1 r 0-1d-0, III p 0- 1d-1d d 1-1-1 r 1d-1d-1d, IV p 0-0-1d d 1-1-1 r 1d-1d; patella I-II d 0-1bristle, III-IV p 1 d 1bristle-1bristle r 1; tibia I p 0-1-0 d 1bristle v 2-2-2ap, II p 1d-1 d 1bristle-1bristle v 2-2-2ap, III-IV p 1-1v d 1-1 r 1-1 v 2-2-2ap; metatarsus I p 0-0-1 d 1bristle-1bristle v 2-2-1 r 0-0-d1, II p 0-0-2 d 2bristle-1bristle r 0-0-1 v 2-2-1ap, III-IV p 1d-1d-2 r 1d-1d-2 v 2-2-1ap.

Measurements: TL 13.22, CL 7.11, CCW 3.67, CTW 5.44, CCH 2.44, CTH 2.56. Eyes: AME 0.36, ALE 0.26, PME 0.65, PLE 0.52, interdistances: AME-AME 0.17; AME-ALE 0.08; PME-PME 0.40; PLE-PLE 1.24. POQ long 1.12, POQ posterior width 1.79, POQ anterior width 1.12. Chelicerae: length 3.19. Opisthosoma: length 6.67, width 4.78. Legs: femur + patella + tibia + metatarsus + tarsus = total length: pedipalp 2.56 + 1.22 + 1.56 + - + 1.78 = 7.12, I 5.00 + 2.78 + 4.00 + 3.78 + 2.11 = 17.67, II 4.89 + 2.33 + 3.78 + 3.78 + 2.22 = 17.09, III 4.33 + 2.11 + 3.11 + 3.72 + 2.11 = 15.38, IV 5.67 + 2.44 + 4.61 + 6.22 + 2.67 = 21.61.

Variation. Females N = 10 (males N = 3) (range, mean±s.d.): TL 10.89 – 18.00, 13.59±1.98; CL 5.67 – 8.17, 6.68±0.69; CW 4.22 – 6.11, 5.02±0.55; (TL 11.67 – 13.33, 12.59±0.69; CL 6.11 – 8.89, 7.26±1.18; CW 4.78 – 6.45, 5.41±0.74).

Distribution. Cuba, Venezuela, Colombia, Peru, Bolivia, Brazil, Paraguay, and Argentina ( Fig. 67 )

Figures

FIGURE 52. Prolycosides aussereri (Keyserling, 1877) comb. nov. Male pedipalp. (MUSM-ENT 500135) a–c (a, retrolateral; b, ventral; c, prolateral); i, stridulatory organ; palp bulb d–h (d, retrolateral; e, ventral; f, prolateral; g, anterior; h, posterior) Scale bars: a–i 200 µm. DMA depression of MA, E embolus, MA median apophysis, P palea, pp pars pendula, S scrapper, ST subtegulum, STA subterminal apophysis, T tegulum, TA terminal apophysis, TDF tibial distal file, TF tegular furrow, TK tegular keel, TL tegular lobe, VP ventral process.

FIGURE 53. Prolycosides aussereri (Keyserling, 1877) comb. nov. Male habitus (MUSM-ENT 500135) a, d, g–h (a, dorsal; d, ventral; g, lateral; h, prosoma, eyes). Female epigynum b, e (b, ventral; e, dorsal). Female habitus c, f, i (c, dorsal; f, ventral; i, lateral) Scale bars: a, c, d, f–i, 1 mm, b, e, 200 µm.

FIGURE 54. Prolycosides aussereri (Keyserling, 1877) comb. nov. Male palpal bulb (MUSM-ENT 500135) a–c (a, ventral; b, palear area; c, MA detail). Scale bars: a, 250 µm, b, 200µm, c, 100 µm. E embolus, MA median apophysis, P palea, pp pars pendula, ST subtegulum, STA subterminal apophysis, T tegulum, TA terminal apophysis, TL tegular lobe, VP ventral process.

FIGURE 55. Prolycosides aussereri (Keyserling, 1877) comb. nov. Female, epigynum (MUSM-ENT 500135) a–c (a, ventral; b, atrium detail; c, copulatory opening). Scale bars: a, 250 µm, b, 200 µm, c, 100 µm. AT atrium, H hood, CO copulatory opening, MS median septum, TS transversal septum.

FIGURE 56. Tarentula aussereri Keyserling, 1877. Female holotype, a. ventral opisthosoma, b. epigynum ventral view. Collecting and ID labels c–d (c, type locality label; d, ID label).

FIGURE 67. Geographic distribution records of Hogna pugil comb. nov. (red cross) updated records taken from Bertkau (1880), Prolycosides aussereri comb. nov. (wine dot).