Centaurea regia var. regia

Negaresh, Kazem & Rahiminejad, Mohammad Reza, 2018, A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae), Phytotaxa 363 (1), pp. 448-450 : 448-450

28a. Centaurea regia var. regia ( Figs. 61 and 62 )

Terminal spine of median appendages (8–) 9–14 mm long; appendages white, sometimes creamish.

Taxonomic and distribution remarks: — Centaurea regia var. regia generally occurs in W Iran, NE and N Iraq, NE Syria and SE Turkey ( Fig. 58 ). The main distribution range of this variety is in Iraq. Centaurea regia var. regia is distinguished from the other varieties by having white or creamish appendages. It has a terminal spine of median phyllaries usually (8–) 9–14 mm long, but there is a few specimens that have a shorter spine (6–8 mm long).

Chromosome numbers: — Centaurea regia var. regia is a diploid taxon with the chromosome number of 2 n = 2 x = 18 ( Fig. 9D and Table 1). This is the first chromosome count for this variety.

Selected specimens examined:— IRAN. Prov. Kermanshah: Shahu mountains , Struass s.n. ( G!) ; 5 km from Ravansar , 1430 m, 14 June 2012, Ranjbar & Negaresh 28868 ( BASU!) ; Ravansar , 1380 m, 14 June 2012, Ranjbar & Negaresh 28828 ( BASU!) ; Ranjbar & Negaresh 28830 ( BASU!) ; Ranjbar & Negaresh 28855 ( BASU!) ; 5 km before bifurcate Ghazel Ghash , 1530 m, 26 May 2012, Ranjbar & Negaresh 28883 ( BASU!) ; Kermanshah toward Javanroud, 25 km before bifurcate Javanroud-Paveh , 1450 m, 30 June 2012, Ranjbar & Negaresh 28893 ( BASU!) ; 5 km to Ravansar, Bave Geh village , 1418 m, Ranjbar & Negaresh 28807 ( BASU!) ; Ranjbar & Negaresh 28808 ( BASU!) ; Kermanshah toward Ravansar, 3 km to Ravansar, 1348 m, 27 May 2014, Negaresh & Kamalnejad 111 (herb. Negaresh !); Negaresh & Kamalnejad 118 (herb. Negaresh !); Ravansar toward Kamyaran , 1–3 km after Ravansar , 1375–1400 m, 29 May 2014, Negaresh & Kamalnejad 20138 ( HUI!) ; Negaresh & Kamalnejad 117 (herb. Negaresh !); Negaresh & Kamalnejad 45547 ( FUMH!) ; Kermanshah toward Ravansar, 1 km to Ravansar, before Sarab-e Jaberi , 34° 41′ 22.3″ N, 46° 40′ 31.2″ E, 1390–1400 m, 23 May 2017, Negaresh 172 (herb. Negaresh!, RUH!) ; Ravansar, benning road of Bashokoui village , 34° 43′ 1.76″ N, 46° 40′ 32.8″ E, 1260–1270 m, 1 June 2015, Negaresh & Asadbegi 182 (herb. Negaresh!, HSB!) . IRAQ. Prov. Arbil: in montibus between Harir and Rawanduz, Bornmüller 1488 ( B!) . Prov. Mosul: 86 km N Mosul toward Zakho , 2 July 1957, Rechinger 10632 ( B!, E!) ; 5 km S Zakho , 700 m, Rawi & Rechinger 16690 ( K!) ; Olaka , 700 m, Rawi 8769 ( K!) . Prov. Kirkuk: near Kirkuk, 27 May 1961, Haines s.n. ( E!) ; Haussknecht s.n. ( G!) . Prov. Sulaymaniyah: roadside bewteen Sheikh Adi and Atush , 700 m, 9 June 1978, Hossain 4269 ( E!, P!: two sheets); Schultz Bipontinus s.n. ( P!) ; Kurdistania Turcica (Assyria) : ad pagum Zacho ( Bidaro , Azach ) dit. Geziret-ibn-Omar , in steppa, 450 m, 6 July 1910, s.c. s.n. ( SAV!) . SYRIA. 1865, Haussknecht s.n. ( P!) . TURKEY. C8 Mardin: between Midyat and Dargeçit , roadside, 20 July 2009, Aslan 3968 ( GAZI!) .

Figures

FIGURE 9. Somatic metaphases of Centaurea spp. A. Centaurea amadanensis subsp. amadanensis (2n = 18). B. C. amadanensis subsp. gymnoclada (2n = 18). C. C. nemecii (2n = 18). D. C. regia var. regia (2n = 18). E. C. regia var. purpurea. Scale A˗E = 2 μm.

FIGURE 58. Distribution of Centaurea regia var. regia.

FIGURE 61. Centaurea regia Boissier var. regia (Kotschy 371, G). A. habit. B. Leaves. C. median part of stem. Scale B and C = 3 cm.

FIGURE 62. Centaurea regia Boissier var. regia (Negaresh 172, herb. Negaresh). A. inflorescence. B. close up of capitule. C. habit and habitat. D. basal leaves. E. taproot and leaves. Scale A = 4 cm, B = 1 cm, C = 15 cm, D–E = 10 cm.