Coremiocnemis Simon 1892

West, Rick C. & Nunn, Steven C., 2010, 2443, Zootaxa 2443, pp. 1-64 : 7-8

Coremiocnemis Simon 1892

( Figs 1–217)

Coremiocnemis Simon 1892b: 146, 1903: 954–956 ; Pocock 1895: 175; Abraham 1924: 1117, pl. 6, figs 27–28; Roewer, 1942: 264; Bonnet, 1959; Raven, 1985: 118, 2005: 18 , figs 1–37; 2002 Raven, Baehr & Harvey (2002: interactive key system); Smith, 1987a: 67, 1987b: 120; Platnick 2009.

Type species: Phlogius cunicularius Simon 1892b by original designation and monotypy.

Etymology: A compound noun whose first element is the Greek noun korema, meaning ‘broom’ or ‘brush’, and whose second element is the Greek feminine noun knemis, meaning ‘shin guard’, which describes the hirsute character of the posterior legs. The gender is feminine.

Diagnosis: Coremiocnemis differs from other Selenocosmiinae genera in having long, recurved setal brushes along retrolateral metatarsus and tarsus IV, in combination with bilobular spermathecae (females), intercheliceral pegs distinct, low cephalothoracic profile, lanceolate, terete embolus emerging anteriorly from the tegulum with distal “kiss curl”, no distal flaring (males). Differs from known Selenocosmia species from Java, Borneo and Sumatra, and all Lyrognathus in the above traits and in the presence of a reduced third claw on tar. IV (except Lyrognathus robustus Smith 1988 ), foveal morphology: strongly procurved foveal groove less in width than the OT (except C. gnathospina sp. nov.), spermathecal morphology: bilobed (all others except C. tropix Raven 2005 : see discussion) with lateral lobes of similar morphology to medials (in those Selenocosmia species with bilobular morphology known to us, lateral lobes display distinctly differing morphology to medials; in Lyrognathus , lateral lobes are only 2/3 length that of medials), anterior metatarsal and tarsal I scopulate morphology: in all Coremiocnemis , anterior scopulae are short, in Selenocosmia species , anterior metatarsal and tarsal scopula are significantly longer/more developed ( Lyrognathus possess similar scopulate morphology to Coremiocnemis in this regard; i.e. short), and in sternal morphology: posterior sternal borders proximally to left and right coxae IV only slightly acuminate (greatly acuminate in both Lyrognathus and Selenocosmia ).

Affinities: Together, Coremiocnemis , Lyrognathus and Indonesian/Malay Selenocosmia share the presence of female bilobular spermathecae in combination with intercheliceral pegs/spines,>3 to 7 retrolateral basomedial cheliceral spines, metatarsus III and/or IV spine arrangement: 1 DD, 2 DV, 1 DPV and 1 DRV, and male lanceolate embolus emerging anteriorly from the tegulum with/without a singular prolateral keel ( West and Nunn 2010). This character combination was not found in any other selencosmiine genus.

Description: Coremiocnemis is a selenocosmiine theraphosid with 15–90 medium to long intercheliceral peg setae, maxillary lyra oval in form, consisting of medium to long shafted paddles with/without distal blades, foveal groove width less than OT (except C. gnathospina sp. nov.), cracked fourth tarsi (except in C. brachyramosa sp. nov.), third claw on tarsi IV, low cephalothoracic profile, retrolateral basomedial cheliceral spines (3 to 7), female with bilobed spermathecae, male with proximally swollen embolus, terete, emerging anteriorly from rounded “popcorn kernel”-shaped tegulum in combination with embolic distal “kiss curl” and no distal flaring, short scopula on anterior metatarsi and tarsi (except C. gnathospina sp. nov.), female with villous recurved retrolateral metatarsi IV setae present or absent, elongated tibiae and metatarsi IV, metatarsi III and/or IV with 1 DD, 2 DV, 1 DPV and 1 DRV spine arrangement, metatarsi IV with scopula divided by thick long setae, scopula only in distal half or less prolaterally, retrolaterally 1/6 to entire. No dorsal carapace thorns or basifemoral thorns.

Species included: Coremiocnemis cunicularia (Simon 1892) , Coremiocnemis brachyramosa sp. nov. West and Nunn, Coremiocnemis gnathospina sp. nov. West and Nunn, Coremiocnemis hoggi sp. nov. West and Nunn, Coremiocnemis jeremyhuffi sp. nov. West and Nunn, Coremiocnemis kotacana sp. nov. West and Nunn, Coremiocnemis obscura sp. nov. West and Nunn, Coremiocnemis valida Pocock 1895 , Coremiocnemis tropix Raven 2005

Distribution: East Malaysia (only Sarawak), State of Brunei Darussalam, Sumatra and West Malaysia. Singapore is removed as part of the range for this genus, the two C. validus Abraham (1924) specimens from Singapore were examined (D. Court, pers. comm.) and do not belong in Coremiocnemis .