Cyana (Gigantovulpecula) detoulgoeti Volynkin, 2020

Volynkin, Anton V., 2020, On the taxonomy of the subgenus Gigantovulpecula Karisch, 2013 of the genus Cyana Walker, 1854, with description of a new species (Lepidoptera: Erebidae Arctiinae: Lithosiini), Zootaxa 4861 (3), pp. 376-384 : 378-379

Cyana (Gigantovulpecula) detoulgoeti Volynkin , sp. n.

( Figs 5, 6 , 13, 14 )

= Chionaema saalmuelleri forme suberythraea Toulgoët, 1971, Bulletin de la Société entomologique de France 76 (2): 85, fig. 6 (Type locality: Madagascar Central , N of Ankazobe, Ambohitantely Forest) .

Type material. Holotype ( Figs 5 , 13 ): male, “ Madagascar C., Ankazobe , Frt. Ambohitanteli, 18–19 II 1955, P. Griveaud–R. Vieu ” / “ PARATYPE ” / “Slide ZSM Arct. 2019- 250♂ A. Volynkin ” (Coll. ZSM).

Paratypes (4 males): 1 male, Madagascar, central area, road from Tananarive to Majunga, 4.IV.[19]65, 80., Collection Jacques Plante / PARATYPE / Chionaema saalmuelleri suberythraea Toulgoët , slide ZSM Arct. 2019- 376 (male) Volynkin (Coll. ZSM); 2 males, EU et P. Griveaud, Tampoketsy, 18.III.53 / Paratypus (Coll. ZSM); 1 male, SE Madagascar, Ranomafana, km. 402, Alt. 650 m, 26.III.–1.IV.55, H. de Toulgoёt (Coll. ZSM).

Remarks. (1) According to the article 45.6.3 of ICZN (1999), the name first published after 1960 as “form” is infrasubspecific, thus, the Toulgoët’s name suberythraea is unavailable. (2) The holotype of Chionaema saalmuelleri forme suberythraea is housed in the MNHN collection (https://science.mnhn.fr/institution/mnhn/collection/el/item/el29068) and remains undissected. According to the articles 72.1.2 and 72.1.3 of ICZN (1999), the holotype and not paratypes has a name-bearing function only, from which it follows that in order for the description of the species to be correct, it is necessary to examine, describe and illustrate the genitalia of the holotype. For this reason, here I designate the dissected male specimen housed in the ZSM collection as holotype. (3) According to the original description ( Toulgoët 1971), besides the holotype, the type series of Chionaema saalmuelleri forme suberythraea includes seven male paratypes from the Ambohitantely Forest being collected by P. Griveaud and R. Vieu in March 1955, and a male paratype from the same locality but collected by C. Moinier on 4.IV.1965. None of the specimens housed in the ZSM collection and labeled as paratypes has such label data therefore these specimens are not paratypes of suberythraea in fact.

Diagnosis. The new species differs clearly from two other members of the subgenus Gigantovulpecula by its relatively small size and intense pale red suffusion on hindwing. The male genitalia of C. detoulgoeti sp. n. are similar to those of C. saalmuelleri , but differ by the more robust distal saccular process, narrower aedeagus with a convex dorsal margin subapically (in C. saalmuelleri that is broader and almost straight), smaller spines on both the dorsal and distal diverticula, and the presence of the 3 rd subdiverticulum in the dorsal diverticulum (whereas in C. saalmuelleri there are two subdiverticula only).

Description. External morphology of adults. Forewing length 14–15 mm in males. Male antenna ciliate. Head creamy white. Thorax ground color creamy white, with crimson spots medially and posteriorly; posterior part of patagia crimson; tegula with crimson sot anteriorly. Abdomen pink medially and creamy whitish basally and apically. Forewing ground color creamy white, transverse lines crimson. Subbasal spot represents large black dot. Subbasal line represents broad but short stroke at costa. Antemedial line nearly straight medially and posteriorly and V-like curved anteriorly. Postmedial line nearly straight, narrowed anteriorly and wavy at costa. Terminal line represents series of various sized spots between veins at wing apex. Cilia creamy white. Hindwing with intense pale red suffusion, but creamy whitish along costa. Cilia creamy whitish. Male genitalia. Uncus short, narrowly trigonal, dorso-ventrally flattened, apically pointed, fully separated from tuba analis. Tuba analis broad, scaphium absent, subscaphium represent broad weakly setose area. Tegumen short and moderately broad. Anellus very broad, heavily scobinated. Juxta broadly X-shaped, moderately sclerotised. Vinculum short, U-shaped, robust. Valva moderately long and broad, broadened medially and narrowed distally, apically rounded. Costa convex medially. Medial crest of valva long but narrow, slightly prominent. Sacculus narrow, its distal process robust, slightly curved dorsally, with short thorn-like tip. Aedeagus moderately long, broadened distally, with convex dorsal wall. Aedeagus vesica broad, curved ventrally; dorsal diverticulum large, bears broad cluster of robust various sized spines and three short but broad subdiverticula; medial diverticulum moderately long, broad, with weakly scobinated area laterally; distal diverticulum short, bears broad and elongate cluster of robust various sized spines.

Female unknown.

Distribution. Central and southeastern Madagascar ( Toulgoët 1971). In southeastern Madagascar (Ranomafa- na) the new species occurs sympatrically with C. saalmuelleri .

Etymology. The species is named after Hervé de Toulgoët, who first described it as ‘ Chionaema saalmuelleri forme suberythraea ’.

Figures

FIGURES 1–8. Cyana spp.: adults. Depositories of specimens: 1–3, 5 and 6 in ZSM; 4 in NHMUK (©); 8 in MWM/ZSM. Figure 7 is given after Hampson (1900).

FIGURES 13–16. Cyana spp.: male (13, 14) and female (15, 16) genitalia. Specimens are deposited in ZSM.