Pseudotibiozus Demange, 1970

Enghoff, Henrik & Larsson, Tobias Berglund, 2018, Pseudotibiozus Demange, 1970 - millipedes of the Eastern Arc Mountains, Tanzania (Diplopoda, Spirostreptida, Spirostreptidae), Zootaxa 4425 (3), pp. 541-554: 542-543

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Pseudotibiozus Demange, 1970


Pseudotibiozus Demange, 1970 

Type species. Mardonius cerasopus Attems, 1914  , by original designation.

Other included species. Pseudotibiozus zophoribates  sp. nov.

Diagnosis. A genus of Spirostreptidae  characterized by the following combination of gonopodal characters: Apical lateral process of metaplica short, rounded. Antetorsal process of telopodite very short, branching off from main telopodite stem before the telopodite curves laterad and exits the coxa. Telopodite becoming thicker shortly after exiting coxa and flattening to an almost ribbonlike structure, telopodite bending mesad at mid-length and dividing into a spinelike branch and a thinner solenomere, solenomere twisting up and around the spinelike branch in an overall mesad direction. Tip of solenomere wrinkled, spongelike.

General description of Pseudotibiozus  .

Males. SIZE. Body length 52–73 mm; diameter 4.4–6.6 mm. 41–43 podous rings, no apodous rings in front of telson.

COLOUR. Live colour contrastful (black body, red legs) in P. cerasopus  ( Fig. 1 View Figure ), not known in P. zophoribates  sp. nov. Metazona of preserved specimens with a light band posteriorly. Dark colouration anteriorly of the band extended as tiny dark 'spike' (ds) into light posterior band at level of ozopore (oz) (even on ozopore-less rings), one or two additional 'spikes' may also be present; 'spikes' difficult to see from the outside ( Fig. 2 C View Figure ), especially on fresh specimens, but conspicuous when the body ring is seen from the inside ( Figs 2 A, B View Figure ).

HEAD ( Figs 3 View Figure , 4 A, B View Figure ). Four supralabral setae. Eyes extending mesad just beyond antennal sockets. Ca. 11 vertical rows of ommatidia, ca. 48 ommatidia. Mandibular stipes with apicoventral lobelike extension. (Details of mandibular gnathal lobe are described under P. cerasopus  ). Gnathochilarium ( Fig. 3 View Figure ): Mentum with median depressed area (mda); lamellae linguales (ll) each with two strong apical setae, one similar-sized seta behind the two and two to three basal setae of varying size; stipites (st) with several setae along latero-apical edge, with or without a field of setae (sbs) basally, with a single stout setae placed in a soft area (ss) near tip.

COLLUM ( Figs 4 A, B View Figure ). Lateral lobes rectangular, with 4–5 striae.

BODY RINGS ( Figs 2 View Figure , 4 View Figure ). Prozonal ring furrows very weak. Metazona vaulted and with longitudinal striae from below ozopores and downward, striae in lover part of striate area more ridgelike and ending in denticles at metazontal hind edge. Ozopore situated ca. 1/3 metazona length behind suture, a weak horizontal suture from ozopore to metazonal hind margin. Sigilla ( Fig. 2 View Figure ) obvious, in one or several rows.

TELSON ( Fig. 4 C View Figure ). Preanal ring with slightly curved dorsal margin, posteriorly with a more or less straight edge, i.e., no proper epiproct; hypoproct a narrow band; anal valves with a submarginal depression, margins forming blunt ‘lips’.

LEGS. As long as body diameter or slightly shorter. Tibia with small apical ventral pad extending under basis of tarsus; no pads on other podomeres. First pair ( Fig. 5 View Figure ) with short, triangular basal prefemoral lobes (pfl), with or without a field of coxosternal setae (cxs).

GONOPODS ( Figs 5 C, D View Figure , 7 View Figure , 8 View Figure ). Sternum (st) blunt triangular to subsemicircular, not reaching as far distal as paracoxites (px). Proplica (pp) straight, ending in a simple apical lobe (apl) with a small patch of short setae at its base. Metaplica (mp) longer than proplica, apically with a triangular-rounded or ovoid lobe (aml). Telopodite (tlp) ( Figs 5 C, D View Figure ) with a very small antetorsal process (atp) originating before the telopodite bends laterad and exits from the coxa, atp hence either completely hidden between pro- and metaplica or barely projecting ( Figs 8 C, D View Figure ). Telopodite ribbonlike from emergence from coxa, ca. at midlength bending mesad and dividing into a mesad spinelike branch (sb) and a thinner solenomere (slm); solenomere twisting around sb in an overall mesad direction. Tip of telopodite strongly wrinkled, spongelike ( Fig. 7 E View Figure ).

Females (known only for P. cerasopus  ). Thicker than males, with relatively shorter legs, lacking modifications of mandibular stipes and legs. 40–44 podous body rings, no apodous rings in front of telson.