Pseudotibiozus cerasopus ( Attems, 1914 )

Enghoff, Henrik & Larsson, Tobias Berglund, 2018, Pseudotibiozus Demange, 1970 - millipedes of the Eastern Arc Mountains, Tanzania (Diplopoda, Spirostreptida, Spirostreptidae), Zootaxa 4425 (3), pp. 541-554: 543-545

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Pseudotibiozus cerasopus ( Attems, 1914 )


Pseudotibiozus cerasopus ( Attems, 1914) 

Figs 1 View Figure , 2 A, C View Figure , 3 A–D View Figure , 4 A View Figure , 5 A, C View Figure , 6 View Figure , 7 View Figure

Mardonius cerasopus Attems, 1914: 139 

Epistreptus cerasopus: Attems (1950) 

Pseudotibiozus cerasopus: Demange (1970) 

Diagnosis. Differs from the only known congener, P. zophoribates  sp. nov., by being smaller (diameter of males 4.4–4.8 mm, vs. 6.0– 6.6 mm), by having the apical proplical lobe almost semi-circular with a short, pointedtriangular or rounded-triangular mesapical process, and by having the apical lobe of the metaplica irregularly triangular.

Material studied (total: 17 ♂♂, 3 ♀♀, 5 juv.)

Holotype: ♂ ZMBAbout ZMB 1368 Ost-Afrika, Conradt leg. ( MfNAbout MfN).

Paratype?: ♂ ZMBAbout ZMB 2050 Buola bei Tanga, Ost-Afrika, Lücker leg 20.vii.1895 ( MfNAbout MfN)

None-type material (All from TANZANIA, all in ZMUCAbout ZMUC): 

East Usambara Mts: 3 ♂♂, 3 juv. Amani, ca. 1000 m asl, 25.vii.–5.viii.1974. I.B. & H. Enghoff leg. (specimens reported by Enghoff & Enghoff 1976; additional data on individual specimens: “forest clearing at the Kwankoro Road”, ”on the loft of a house”, “on road by night”, “Amani W Forest Reserve, at Dodwe River”, “Amani Pond, under dry bark”); 1 ♂ Amani, ca. 1000 m asl, 1.ix.1981, N. Scharff & M. Stoltze leg.

Kanga Mts : 1 ♂ 400 m asl, 23.xi.1984, M. Stoltze leg.  ; 1 ♀ Morogoro Region, Kanga Forest Reserve, 400-500 m asl, 22-25.xi.1984, lowland rain forest, N. Scharff leg. 

Udzungwa Mts: 1 ♂, 1 juv. Mwanihana Forest above Sanje , 1000m asl, 1.viii.1982, M. Stoltze & N. Scharff leg.  ; 1 juv. Mwanihana Forest, Sanje River , 1400 m asl, litter, 15.xii.1982, M. Stoltze & N. Scharff leg.  ; 2 ♂♂ Morogoro Region, Udzungwa Mts National Park, Mizimu Camp , 1.ix.2013, 07°48’23.40”S, 36°51’7.29”E, 769 m. Hand collected. T. Pape & N. Scharff leg.GoogleMaps  ; 4 ♂♂ Morogoro Region, Kilombero District, Udzungwa Mts National Park, Mizimu Camp , 19-21.viii. 2017, 780 m asl, 07°48.390’S, 36°51.123E. Hand collected in forest, night collecting, J. Pedersen leg.GoogleMaps  ; 3 ♂♂ Morogoro Region, Udzungwa Mts National Park, Sanje Chini camp, 598m, 07°46’24.6”S, 36°54’.47.7”E, 17-19.i.2014, hand collected. T. Pape & N. Scharff leg.  ; 1 ♀ Morogoro Region, Udzungwa Mts National Park, Mizimu Camp , 3.ix.2012, 07°48’23.40”S, 36°51’7.29”E, 769 m, hand collected. T. Pape & N. Scharff leg.GoogleMaps  ; 1 ♀ Morogoro Region, Udzungwa Mts National Park, trail to Mizimu Camp , 250 m asl., hand collected 3.ix.2012. T. Pape & N. Scharff leg. 

Description. Males.

SIZE. Body length 52–60 mm. Diameter 4.4–4.8 mm. 41–43 body rings.

COLOUR. Live colour ( Fig. 1 View Figure ) black, prozonites dull with a greyish tinge, metazonites shining deep black; antennae red-brown; legs contrastingly bright red; superficially resembling a miniature version of the common East African Epibolus pulchripes ( Gerstäcker, 1873)  (fam. Pachybolidae  , order Spirobolida  ). Colour of preserved specimens faded, legs reddish orange.

MANDIBLES ( Fig. 6 View Figure ). Stipes with apicoventral lobe, a soft pad apically on the lobe. Odontomere (od) fused with psectromere (ps); sectile edge (se) of psectromere with four lobes; a meso-dorsal lobe (mdl) present between od and se. 11 pectinate lamellae (pl). Molar plate with a distal transverse groove (mg) occupying ca. 1/6 of the mesal surface. Dorsal surface of mandible with a group of small pointed pegs (pp) between pectinate lamellae and molar plate ( Fig. 6 D View Figure ).

GNATHOCHILARIUM ( Figs 3 A–D View Figure ). Basal edge of median depression of mentum (mda) sharp, overlying main part of mentum and thus delimiting an arched, transverse cavity. Stipites (st) with a longitudinal patch of setae (sbs) in basal half.

BODY RINGS ( Fig. 2 A View Figure ). A single row of large sigilla; sometimes additional, incomplete rows of smaller sigilla.

LEGS. As long as body diameter. First pair with a patch of setae on coxosternum ( Fig. 5 A View Figure ).

GONOPODS ( Figs 5 C View Figure , 7 View Figure ). Proplica parallel-sided, apically with an almost semi-circular lateral lobe (apl), mesapically with a short, triangular process (map). Metaplica with mesal margin turning laterad at ca. 45° just before end of proplica, apical lobe (aml) subtriangular, projecting laterad and apicad, more or less resembling the head of a seal. A sub-circular sport of dark colour subbasally on posterior surface of metaplica, position of dark spot variying between individuals, but always symmetrical. Antetorsal process (atp) of telopodite ( Fig. 5 C View Figure ) projecting distad, completely hidden between pro- and metaplica.

Females. SIZE. Body length 57–61 mm. Diameter 5.4–5.7 mm. 40–44 body rings.

LEGS. Length 0.75 × body diameter.

Distribution. Known from three massifs of the Eastern Arc Mountains: East Usambara, Kanga and Udzungwa Mts ( Fig. 9 View Figure ).


Museum f�r Naturkunde Berlin (Zoological Collections)


Museum f�r Naturkunde


Zoological Museum, University of Copenhagen














Pseudotibiozus cerasopus ( Attems, 1914 )

Enghoff, Henrik & Larsson, Tobias Berglund 2018


Mardonius cerasopus

Attems, 1914 : 139