Pseudotibiozus zophoribates

Enghoff, Henrik & Larsson, Tobias Berglund, 2018, Pseudotibiozus Demange, 1970 - millipedes of the Eastern Arc Mountains, Tanzania (Diplopoda, Spirostreptida, Spirostreptidae), Zootaxa 4425 (3), pp. 541-554: 545-552

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Pseudotibiozus zophoribates

sp. nov.

Pseudotibiozus zophoribates  sp. nov.

Figs 2 BView FIGURE 2, 3 EView FIGURE 3, 4 B, CView FIGURE 4, 5 B, DView FIGURE 5, 8View FIGURE 8

Diagnosis. Differs from the only known congener, P. cerasopus  , by being larger (diameter of males 6.0– 6.6 mm, vs. 4.4–4.8 mm), by having the apical proplical lobe almost semi-circular with a parallel-sided, broadly or rounded mesapical process, and by having the apical lobe of metaplica almost egg-shaped.

Etymology. The name is a composite Greek noun in apposition, from zophos (darkness, dusk, also referring to the West) and oribates (mountaineer); it refers to the type locality, the West Usambara Mountains.

Material studied (total: 2 ♂♂)

Holotype: 1 ♂ TANZANIA, Tanga, W Usambara Mts, Mazumbai forest , 4°49’S, 38°30’E, 1400–1800 m asl, 10–20.xi.1995, C.E. Griswold, N. Scharff & D. Ubick leg. ( ZMUCAbout ZMUC)GoogleMaps 

Paratype: 1 ♂ same data ( ZMUCAbout ZMUC). 

Description. Males.

SIZE. Body length 71–73 mm. Diameter 6.0– 6.6 mm. 43 body rings.

COLOUR (preserved specimens). Antennae and legs red, although less intensely so than in preserved specimens of P. cerasopus  . Prozona and anterior part of metazona yellowish brown, posterior part of metazona dark brown, semi-transparent edge of metazona reddish.

MANDIBLES. Stipes with apicoventral lobe, no soft pad apically on the lobe. Gnathal lobe not studied with SEM, but apparently no group of small pointed pegs between pectinate lamellae and molar plate on dorsal surface, and no meso-dorsal lobe present between odontomere and sectile edge of psectromere.

GNATHOCHILARIUM ( Fig. 3 EView FIGURE 3). Basal edge of median depression of mentum blunt, not overlying main part of mentum, not delimiting a proper cavity. Stipites without setae in basal half.

BODY RINGS ( Fig. 2 BView FIGURE 2). Several rows of sigilla; rows sometimes irregular, especially below ozopore level.

LEGS. Length 0.9 × body diameter. First pair with only a few setae on coxosternum, not a large patch as in P. cerasopus  . ( Fig. 5 BView FIGURE 5).

GONOPODS ( Figs 5 DView FIGURE 5, 8View FIGURE 8). Proplica club-shaped: base relatively narrow, largest width subapically. Apical lateral lobe (apl) with regularly rounded margin continuing in broadly rounded mesapical lobe. Metaplica with mesal margin turning laterad at ca. 70° just before end of proplica, forming a blunt “corner”; apical lobe (aml) tongue- or egg-shaped, hardly projecting laterad. Antetorsal process (atp) of telopodite ( Fig. 5 DView FIGURE 5) projecting obliquely mesodistad, tip just visible between pro- and metaplica ( Figs 8 C, DView FIGURE 8).

Females. Unknown.

Distribution. Only known from the type locality, Mazumbai forest in the West Usambara Mountains ( Fig. 9View FIGURE 9).


Zoological Museum, University of Copenhagen