Progyrolepis FRITSCH , 1895

Štamberg, Stanislav, 2018, Actinopterygians Of The Permian Locality Buxières-Les-Mines (Bourbon- L’Archambault Basin, France) And Their Relationship To Other Early Actinopterygians, Fossil Imprint 74 (3 - 4), pp. 245-291 : 248

Genus Progyrolepis FRITSCH, 1895

T y p e s p e c i e s. Progyrolepis speciosus ( FRIČ, 1875)

I n c l u d e d s p e c i e s. Progyrolepis speciosus ( FRIČ,

1875); P. heyleri POPLIN, 1999

G e o l o g i c a n d g e o g r a p h i c d i s t r i b u t i o n. Upper Carboniferous – Lower Permian, Bourbonl’Archambault Basin ( France), Puertollano Basin ( Spain); Central and West Bohemian Late Palaeozoic basins and Krkonoše Piedmont Basin (the Czech Republic).

D i a g n o s i s. After Štamberg (1991; here modified). Carnivorous fish reaching 60–70 cm in total length. Anterior lepidiotrichia of the pectoral fin unjointed proximally, distally articulated and dichotomously branched. Endocranium ossified. Frontal is 2.5 times longer than wide with sculpture formed by tubercles. Rostral region of the head is conspicuously convex anteriorly, comprising a large single postrostral, paired nasal and denticulated rostropremaxilla. The medial side of the palatoquadrate is formed by the entopterygoid, ectopterygoid, dermometapterygoid and small dermopalatines. The maxilla has a long maxillary plate that is trapezoidal in shape with an outstanding process ventroposteriorly. The length – height ratio of the maxillary plate is 1.6–2. The length of the maxilla is 1.6 to 1.9 times that of the maxillary plate length. The maxillary plate bears outstanding sculpture formed by vermicularly curved ridges. Tubercles prevail along the ventral border and on the anterior suborbital part of the maxilla. The large anterior region of the maxillary plate is without sculpture and is overlapped by a large jugal. The lower jaw is very stout. The teeth on the jaws are arranged in two rows in which the outer row consists of numerous small sharply pointed teeth whereas a small number of large conical laniaries form the inner row. The preoperculum is conspicuously bent at an angle of 130°, and inclined anteriorly. The orbit is small and located anteriorly. The operculum is of oval or oblong shape, three times deeper than it is long and 2.5 times deeper than the depth of the quadrate suboperculum. An antoperculum or epipraeoperculum may be present. Numerous branchiostegal rays. Stout rhomboidal scales with conspicuous sculpture formed by ridges which diagonally traverse across the scale and terminate in denticulation posteriorly.